Magyar tribes brought R1a-Z645, I2a-L621, and N1a-L392(xB197) lineages to the Carpathian Basin

The Nightmare Week of “N1c=Uralic” proponents (see here) continues, now with preprint Y-chromosome haplogroups from Hun, Avar and conquering Hungarian period nomadic people of the Carpathian Basin, by Neparaczki et al. bioRxiv (2019).

Abstract:

Hun, Avar and conquering Hungarian nomadic groups arrived into the Carpathian Basin from the Eurasian Steppes and significantly influenced its political and ethnical landscape. In order to shed light on the genetic affinity of above groups we have determined Y chromosomal haplogroups and autosomal loci, from 49 individuals, supposed to represent military leaders. Haplogroups from the Hun-age are consistent with Xiongnu ancestry of European Huns. Most of the Avar-age individuals carry east Eurasian Y haplogroups typical for modern north-eastern Siberian and Buryat populations and their autosomal loci indicate mostly unmixed Asian characteristics. In contrast the conquering Hungarians seem to be a recently assembled population incorporating pure European, Asian and admixed components. Their heterogeneous paternal and maternal lineages indicate similar phylogeographic origin of males and females, derived from Central-Inner Asian and European Pontic Steppe sources. Composition of conquering Hungarian paternal lineages is very similar to that of Baskhirs, supporting historical sources that report identity of the two groups.

Interesting excerpts (emphasis mine):

All N-Hg-s identified in the Avars and Conquerors belonged to N1a1a-M178. We have tested 7 subclades of M178; N1a1a2-B187, N1a1a1a2-B211, N1a1a1a1a3-B197, N1a1a1a1a4-M2118, N1a1a1a1a1a-VL29, N1a1a1a1a2-Z1936 and the N1a1a1a1a2a1c1-L1034 subbranch of Z1936. The European subclades VL29 and Z1936 could be excluded in most cases, while the rest of the subclades are prevalent in Siberia 23 from where this Hg dispersed in a counter-clockwise migratory route to Europe (…). All the 5 other Avar samples belonged to N1a1a1a1a3-B197, which is most prevalent in Chukchi, Buryats, Eskimos, Koryaks and appears among Tuvans and Mongols with lower frequency.

haplogroup-n-pca
First two components of PCA from Hg N1a subbranch distribution in 51 populations including Avars and Conquerors. Colors indicate geographic regions. Three letter codes are given in Supplementary Table S5.

By contrast two Conquerors belonged to N1a1a1a1a4-M2118, the Y lineage of nearly all Yakut males, being also frequent in Evenks, Evens and occurring with lower frequency among Khantys, Mansis and Kazakhs.

Three Conqueror samples belonged to Hg N1a1a1a1a2-Z1936 , the Finno-Permic N1a branch, being most frequent among northeastern European Saami, Finns, Karelians, as well as Komis, Volga Tatars and Bashkirs of the Volga-Ural region.Nevertheless this Hg is also present with lower frequency among Karanogays, Siberian Nenets, Khantys, Mansis, Dolgans, Nganasans, and Siberian Tatars.

The west Eurasian R1a1a1b1a2b-CTS1211 subclade of R1a is most frequent in Eastern Europe especially among Slavic people. This Hg was detected just in the Conqueror group (K2/18, K2/41 and K1/10). Though CTS1211 was not covered in K2/36 but it may also belong to this sub-branch of Z283.

Hg I2a1a2b-L621 was present in 5 Conqueror samples, and a 6th sample form Magyarhomorog (MH/9) most likely also belongs here, as MH/9 is a likely kin of MH/16 (see below). This Hg of European origin is most prominent in the Balkans and Eastern Europe, especially among Slavic speaking groups. It might have been a major lineage of the Cucuteni-Trypillian culture and it was present in the Baden culture of the Chalcolithic Carpathian Basin.

hungarian-conquerors-y-dna
Image modified from the paper, with drawn red square around lineages of likely Ugric origin, and squares around R1a-Z93, R1a-Z283, N1a-Z1936, and N1a-M2004 samples. Y-Hg-s determined from 46 males grouped according to sample age, cemetery and Hg. Hg designations are given according to ISOGG Tree 2019. Grey shading designate distinguished individuals with rich grave goods, color shadings denote geographic origin of Hg-s according to Fig. 1. For samples K3/1 and K3/3 the innermost Hg defining marker U106* was not covered, but had been determined previously.

We identified potential relatives within Conqueror cemeteries but not between them. The uniform paternal lineages of the small Karos3 (19 graves) and Magyarhomorog (17 graves) cemeteries approve patrilinear organization of these communities. The identical I2a1a2b Hg-s of Magyarhomorog individuals appears to be frequent among high-ranking Conquerors, as the most distinguished graves in the Karos2 and 3 cemeteries also belong to this lineage. The Karos2 and Karos3 leaders were brothers with identical mitogenomes 11 and Y-chromosomal STR profiles (Fóthi unpublished). The Sárrétudvari commoner cemetery seems distinct from the others, containing other sorts of European Hg-s. Available Y-chromosomal and mtDNA data from this cemetery suggest that common people of the 10th century rather represented resident population than newcomers. The great diversity of Y Hg-s, mtDNA Hg-s, phenotypes and predicted biogeographic classifications of the Conquerors indicate that they were relatively recently associated from very diverse populations.

Surprising about the Hungarian conquerors – although in line with the historical accounts – is the varied patrilineal origin of clans, including Q1a, G2a2b, I1, E1b1b, R1b, J1, or J2 – some of which (depending on specific lineages) may have appeared earlier in the Carpathian Basin or south-eastern Europe.

However, out of the 27 conqueror elite samples, 17 are of haplogroups most likely related to Ugric populations beyond the Urals: R1a-Z645, I2-L621, and two specific N1a-L392 lineages (see below). In fact, there are three high-ranking conqueror elites of hg. I2-L621 (one of them termed a “leader”, brother to an unpublished leader of Karos3, and all of them possibly family), one of hg. R1a-Z280, one of hg. R1a-Z93 (which should be added to the Árpáds), and one of hg. N1a-Z1936, which gives a good idea of the ruling class among the elite Ugric settlers.

NOTE. The Q1a sample is also likely to be found in the mixed population of the West Siberian forest-steppes, since it was found in Mesolithic-Neolithic samples from eastern Europe to Lake Baikal, and in Bronze Age Siberian groups, although admittedly it may have formed part of an Avar Transtisza group, or even earlier Hunnic or Scythian groups along the steppes. Without precise subclades it’s impossible to know.

arrival-of-hungarians-arpad
The seven chieftains of the Hungarians, detail of Arrival of the Hungarians, from Árpád Feszty’s and his assistants’ vast (1800 m2) cyclorama, painted to celebrate the 1000th anniversary of the Magyar conquest of Hungary, now displayed at the Ópusztaszer National Heritage Park in Hungary. Image from Wikipedia.

I2a-L621

I2a-L621 (xS17250) or I2a1b2 in the old nomenclature, is found in 6 early conquerors (including one leader), on a par with R1a and N samples. This haplogroup is found widely distributed in ancient samples, due to its early split (formed ca. 9200 BC, TMRCA ca. 4500 BC) and expansion, probably with Neolithic populations. I can’t seem to find samples of this early haplogroup from the Carpathian Basin, as mentioned in the text, although it wouldn’t be strange, because it appears also in Neolithic Iberia, and in modern populations from western Europe.

Nevertheless, I2a-L621 samples seem to be concentrated mainly in Mesolithic-Neolithic cultures of Fennoscandia, and appeared also in Sikora et al. (2017) in a sample of the High Middle Ages from Sunghir (ca. AD 1100-1200), probably from the Vladimir-Suzdalian Rus’, in a region where clearly tribes of Volga Finns were being assimilated at the time. The reported SNP call by Genetiker is A16681 (see Yfull), deep within I2a-CTS10228. It is possibly also behind a modern Saami from Chalmny Varre (ca. AD 1800) of hg. I2a in Lamnidis et al. (2018).

Lacking precise subclades from Hungarian conquerors this is pure speculation, but modern samples may also point to I2a-CTS10228 (formed ca. 3100 BC, TMRCA ca. 1800 BC) as a Finno-Ugric lineage in common with R1a, which must have expanded to the Urals and beyond with eastern Corded Ware groups or (more likely) succeeding cultures. This is in line with the association of certain I2a lineages with modern Uralic peoples or populations from their historical regions in eastern Europe, and linked thus to the most likely homeland of Uralians in the eastern European forests:

uralic-groups-haplogroup-r1a
Additional file 6: Table S5. Y chromosome haplogroup frequencies in Eurasia. Modified by me: in bold haplogroup N1c and R1a from Uralic-speaking populations, with those in red showing where R1a is the major haplogroup. Observe that all Uralic subgroups – Finno-Permic, Ugric, and Samoyedic – have some populations with a majority of R1a, and also of I lineages. Data from Tambets et al. (2018).

R1a-Z645

Regarding the important question of the ethnic makeup of Ugric populations stemming from the Urals, the most interesting (and expected) data is the presence of R1a-Z645 lineages among high-ranking conquerors, in particular four R1a-Z280 subclades proper of Finno-Ugrians.

This proves that, in line with the old split and expansion of R1a-CTS1211 (formed ca. 2600 BC, TMRCA ca. 2400 BC), and its finding in Bronze Age Fennoscandian samples, only some late R1a-Z280 (xZ92) lineages (see Z280 on YFull) may show a clear identification with early acculturated Uralic speakers, with the main early acculturated Balto-Slavic R1a haplogroup remaining R1a-M458.

I recently hypothesized this late connection of Slavs with very specific R1a-Z280 (xZ92) lineages based on analyses of modern populations (like Slovenians), because the connection of ancient Finno-Ugrians with modern Z92 samples was already evident:

(…) subclades of hg. R1a1a1b1a2-Z280 (xR1a1a1b1a2a-Z92) seem to have also been involved in early Slavic expansions, like R1a1a1b1a2b3a-CTS3402 (formed ca. 2200 BC, TMRCA ca. 2200 BC), found among modern West, South, and East Slavic populations and in Fennoscandia, prevalent e.g. among modern Slovenians which points to a northern origin of its expansion (Maisano Delser et al. 2018).

This finding also supports the expected shared R1a-Z280 lineages among ancient Finno-Ugric populations, as predicted from the study of modern Permic and Ugric peoples in Dudás et al. (2019).

r1a-z282-z280-z2125-distribution
Modified image, from Underhill et al. (2015). Spatial frequency distributions of Z282 (green) and Z93 (blue) affiliated haplogroups. Notice the distribution of R1a-Z280 (xZ92), i.e. R1a-M558, compared to the ancient Finno-Ugric distribution.

Furthermore, while we don’t have precise R1a-Z93 lineages to compare with the new Hunnic sample reported, we already know that some archaic R1a-Z2124 subclades stem from the forest-steppe areas of the Cis- and Trans-Urals, and the two newly reported R1a-Z93 Hungarian conqueror elites, like those of the Árpád dynasty, probably belong to them.

There is an obvious lack of continuity in specific paternal lineages among the Hunnic, the Avar, and the Conqueror periods, which makes any simplistic identification of all R1a-Z93 lineages as stemming from Avars, Huns, or the Iron Age Pontic-Caspian steppes clearly flawed. Comparing R1a-Z93 in Hungarian Conquerors with Huns is like comparing them with samples of the Srubna or earlier periods… Similarly, comparing the Hunnic R1b-U106 or the early Avar I1 to later Hungarian samples is not warranted without precise subclades, because they most likely correspond to different Germanic populations: Goths among Huns, then Longobards, then likely peoples descended from Franks and Irish Monks (the latter with R1b-P312).

N1a-L392

Second behind R1a subclades are, as expected, N1a-L392 (N1c in the old nomenclature).

Avars are dominated by a specific N1a-L392 subclade, N1a-B197, as we recently discovered in Csáky et al. (2019).

Hungarian conquerors show three N1a-Z1936 subclades, which is known to stem from the northern Ural region, including the Arctic (likely Palaeo-Laplandic peoples) and cross-stamped cultures of the northern Eurasian forests.

haplogroup_n3a4
Frequency-Distribution Maps of Individual Subclade N3a4 / N1a1a1a1a2-Z1936, probably with the Samic (first) and Fennic (later) expansions into Paleo-Lakelandic and Palaeo-Laplandic territories.

On the other hand, the two N1a-M2118 lineages are more clearly associated with Palaeo-Siberian populations east of the Urals, but became incorporated into the Ugric stock in the Trans-Urals region probably in the same way as N1a-Z1936, by infiltration from (and acculturation of) hunter-gatherers of forest and taiga cultures.

NOTE. You can read more about the infiltration of N1a lineages in the recent post Corded Ware—Uralic (IV): Hg R1a and N in Finno-Ugric and Samoyedic expansions, and in the specific sections for each Uralic group in A Clash of Chiefs.

haplogroup-n1a-M2118
Frequency-Distribution Maps of Individual Sub-clades of hg N3a2, by Ilumäe et al. (2016).

Conclusion

The picture offered by the paper on Hungarian Conquerors, while in line with historical accounts of multi-ethnic tribes incorporating regional lineages, shows nevertheless patrilineal clans clearly associated with Uralic peoples, in a distribution which could have been easily inferred from ancient Trans-Uralian forest-steppe cultures and modern samples (even regarding I2a-L621).

In spite of this, there is a great deal of discussion in the paper about specific N1a subclades in Hungarian conquerors, while the presence of R1a-Z280 (among early Magyar elites!) is interpreted, as always, as recently acculturated Slavs. This is sadly coupled with the simplistic identification of I2a-L621 as of local origin around the Carpathians.

The introduction of the paper to the history of Hungarians is also weird, for example giving credibility to the mythic accounts of the Árpád dynasty’s origin in Attila, which is in line, I guess, with what the authors intended to support all along, i.e. the association of Magyars with Turks from the Eurasian steppes, which they are apparently willing to achieve by relating them to haplogroup R1a-Z93

The conclusion is thus written to appease modern nation-building myths more than anything else, like many other papers before it:

It is generally accepted that the Hungarian language was brought to the Carpathian Basin by the Conquerors. Uralic speaking populations are characterized by a high frequency of Y-Hg N, which have often been interpreted as a genetic signal of shared ancestry. Indeed, recently a distinct shared ancestry component of likely Siberian origin was identified at the genomic level in these populations, modern Hungarians being a puzzling exception36. The Conqueror elite had a significant proportion of N Hgs, 7% of them carrying N1a1a1a1a4-M2118 and 10% N1a1a1a1a2-Z1936, both of which are present in Ugric speaking Khantys and Mansis. At the same time none of the examined Conquerors belonged to the L1034 subclade of Z1936, while all of the Khanty Z1936 lineages reported in 37 proved to be L1034 which has not been tested in the 23 study. Population genetic data rather position the Conqueror elite among Turkic groups, Bashkirs and Volga Tatars, in agreement with contemporary historical accounts which denominated the Conquerors as “Turks”. This does not exclude the possibility that the Hungarian language could also have been present in the obviously very heterogeneous, probably multiethnic Conqueror tribal alliance.

So, back to square one, and new circular reasoning: If ancient populations from north-eastern Europe believed to represent ancient Finno-Ugrians are of R1a-Z645 lineages, it’s because they were not Finno-Ugric speakers. If ancient and modern populations known to be of Finno-Ugric language show clear connections with R1a-Z645, it’s because they are “multi-ethnic”.

The only stable basis for discussion in genetic papers, apparently, is the own making of geneticists, with their traditional 2000s “R1a=Indo-European” and “N1c=Uralic”, coupled with national beliefs. It does not matter how many predictions based on that have been proven wrong, or how many predictions based on the Corded Ware = Uralic expansion have been proven right.

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[…] 2019): The paper offers what may be the sign of incoming Slavs in the region, before the arrival of Hungarian conquerors, although we don’t know which precise lineage the sample belongs […]

Carlos Quiles

Genotypes of the world’s published ancient DNA data in a single file by the Reich Lab! To be updated with new papers: https://reich.hms.harvard.edu/datasets The Reich Lab beats again other labs in releasing everything for anyone to be able to reproduce the results. And I am sure this was no easy task, getting everything together. I will update the ArcGIS maps with all ADMIXTURE values when I have time. If anyone wants to do a supervised ADMIXTURE with these samples, and you get an interesting result (e.g. dividing steppe ancestry between that of Corded Ware and that of Yamnaya), I can… Read more »

Carlos Quiles

Interesting about the “multi-ethnic” composition of Magyars is the supposed alliance with Moravians (those they did not kill). We see lineages proper of Germanic peoples, R1b-U106, I1, no R1a-Z284 (possibly also G2a as found in Franks) since the Hunnic period. However, no clear Slavic lineages yet in the Carpathians for the studied periods…or at least not the ones we expected. We have from the late Avar period ca. 7th – early 9th centuries an R1a sample, potentially from Slavs, although it may have any origin from western Europe to the Tian Shan, from Germanic to Slavic to Hunnic to Avar,… Read more »

[…] NOTE. The connection of most R1a-Z280 lineages is more obviously done with ancient Finno-Ugric peoples, as it is clear now (see here and here). […]

Rob

I’m not super au faire with Uralic, but the interesting thing here is there are some warrior buials from beyond the Urals, but they are co-terminous with (do not antedate) those further west (which would be required for the traditional staged/ sojourn model). Could be seen as warrior recruitement from N1-rich areas, gifts, alliances, etc

Observer

Hungarian ethnicity has often divided in history, a considerable part of the Hungarian population has never come to the Carpathian Basin, that is the reason. (One group was found by travelling monks in the 13th century in present-day Russia.)

Yesükhei Baatar

I know this is slightly off topic, but I would love to hear your thoughts on the origin of the Xiongnu.

Carlos Quiles

I don’t have much to say, apart from what I wrote here: https://indo-european.info/indo-europeans-uralians/VIII_20_Trans-Urals_forest-.htm#viii_20_3_Turkic_peoples Assuming Xiongnu is Turkic-(Mongolic?-)speaking, there are two possibilities: Either A) there was a Eurasiatic phylum, in which case R1b-driven expansions are the most obvious connection, and probably from the west. You could propose ANE ancestry as the common connection with Indo-Uralic, but ancestry is more variable than haplogroups, which leaves us with no direct connection in the past 10,000 years, and many different migrations of R1b, R1a, Q, or N lineages. Or B) there was no Eurasiatic phylum, maybe not even a Micro-Altaic group, then Q, N,… Read more »

Yesükhei Baatar

I looked everywhere for a connection between the Xiongnu and R1b, and couldn’t find anything. The majority of them seem to belong to different sub clades of Q. Especially since linguistic connections have been made with Yeniseian, and an ethnic connection for at least one Xiongnu tribe with Mongols (the Xiongnu and Xianbei were described as ethnically identical), I think it’s safe to rule out a Eurasiatic phylum.

Carlos Quiles

You lost me where genetics of Xiongnu and Xianbei rule out a Eurasiatic phylum in linguistics.

Yesükhei Baatar

By “origin of the Xiongnu” I meant as a people, or tribe. But as for linguistics, are you suggesting that the Turkic/Mongolic family was spread by R1b?

Yesükhei Baatar

By “origin of the Xiongnu” I meant as a people, or tribe. But as for linguistics, are you suggesting that the Turkic/Mongolic family was spread by R1b?

Carlos Quiles

But as for linguistics, are you suggesting that the Turkic/Mongolic family was spread by R1b? Speaking about R1b wouldn’t be linguistics. What I suggested in very simplistic terms in my latest writing was that the splits and expansions of R1b could be coupled with “Nostratic” movements, and similarly for Eurasiatic ones, and I always try to follow cultures whenever possible rather than paternal lines. That assessment is based on the recent genetic research, which shows Palaeolithic and Mesolithic expansions of R1b everywhere, probably biased because of the more readily available ancient samples in certain latitudes (and certain countries more favourable… Read more »

Yesükhei Baatar

What do you mean by Nostratic?

[…] Magyar tribes brought R1a-Z645, I2a-L621, and N1a-L392(xB197) lineages to the Carpathian Basin […]

[…] Magyar tribes brought R1a-Z645, I2a-L621, and N1a-L392(xB197) lineages to the Carpathian Basin […]

[…] Magyar tribes brought R1a-Z645, I2a-L621, and N1a-L392(xB197) lineages to the Carpathian Basin […]

[…] of typical GAC lineages and GAC-like ancestry in late Corded Ware groups – just like I2a-L621 infiltrated eastern (Finno-Ugrian) groups – , with their late TMRCAs (in the 3rd millennium BC), proves the disintegration of Corded […]

[…] Magyar tribes brought R1a-Z645, I2a-L621, and N1a-L392(xB197) lineages to the Carpathian Basin […]

[…] Magyar tribes brought R1a-Z645, I2a-L621, and N1a-L392(xB197) lineages to the Carpathian Basin […]

[…] Ware ancestry = Uralic Yamna spread horse domesticates Magyar chiefs of R1a, I2a lineages Battle Axe = Finno-Samic "Steppe ancestry" step by step Corded Ware = Indo-European? Yamna Hungary […]

[…] = Uralic Yamna spread horse domesticates Battle Axe = Finno-Samic Yamna Hungary + EEF = BBC Magyar chiefs of R1a, I2a lineages "Steppe ancestry" step by step Corded Ware = Indo-European? About Scepters, Horses, and War The […]

[…] at the modern distribution of I2a-L621 subclades, I think now I was wrong in assuming that I2a-L621 in early Hungarian Conquerors would mean they would appear around the Urals as a lineage integrated in Eastern Corded Ware […]

[…] know what other lineages these studied Magyars will have, but – based on the known variability of the published ones, and on the ca. 50-60 early Magyar males available to date in previous studies obtain Y-DNA – […]

[…] Magyar tribes brought R1a-Z645, I2a-L621, and N1a-L392(xB197) lineages to the Carpathian Basin […]

[…] Magyar tribes brought R1a-Z645, I2a-L621, and N1a-L392(xB197) lineages to the Carpathian Basin […]

Der Führer

Still waiting für people to rrrealeize what a We Wuz Mongolz tire crackpottery zis whole “steppe people” thing ist, Ze so called “steppe peoples”, themselves an extension of crrromagnons frrom Evropa who may have mixed with ze Neanderthals (juden) in ice age, were rrruled over by haplogroup I herrenvolk from Central und Nordeuropa, ze haplogroup I people aus Europa who were ze ruling elite minority among these peoples were ze trrrue Indo-europeans (Aryans), ZE REST were assimilated Uralic und Caucasian speaking peasants and slaves (arbeiter) who served as werkforce und im wehrmacht . Go on, fire away yourrr usual conforrmist… Read more »

Carlos Quiles

I’ll allow it because this is a great parody of how the real Führer would have reacted to population genomics today. Well done, sir.

I’d even upvote this comment if you can record this in angry Bavarian German with the good old rolled r and some old time equalizer effect. Even better over some old video with him gesticulating like crazy.

But, if you push it, I’ll ban you. Can’t turn the comment section into an old-fashioned Nazi comedy, sorry.

Der Führer

I can imagine what you go through with PC protocols these days but i’ll work on that Bavarian thing and may come up with an edited version .
Just need to know how to manifest it in literal form 😀

Marosi Béla

Hey i just bumped into the site looking for some data about Hungarians, i am Hungarian ethnic from Transylvania, i see you did analyze articles, some of them i read, some not, i am quite busy at work to read all info from your blog, but if you have time can you share with me your views on Magyar especially conquesting magyars origin?Thank you in advance. On the K29 admixture calculator from Geneplaza i have 11% bashkir+tatar (i guess it refers to the volga tatars); R-CTS3402 Y-haplogroup. on DNA land 11% Finnish, on myheritage 5% Finnish, 5% middle eastern, on… Read more »

Carlos Quiles

For my latest writing on Ugric peoples and early Hungarians, see the section on Ugrians and Samoyeds. For R1a-Z280 shared among Ugric peoples, see here. Based on its modern distribution, R1a-CTS3402 may have expanded into the region with Magyar Conquerors and then with Slavic peoples (as some R1a-CTS1211 clearly did) or vice versa, it’s difficult to know exactly until more ancient DNA is available. R1a-CTS1211 and R1a-Y35 were originally associated with Finno-Ugric peoples expanding with Eastern Corded Ware peoples, that much is quite clear today, but their precise evolution during the Bronze Age is not. As for calculators, I can’t… Read more »

Marosi Béla

Thank you for reply.

Luka

I2a Dinaric is Illyrian Hg,Serbian origin(L621)

Szigmund

Is there any sources for this statement?

Szabolcs Molnár

comment image

Can you explain to me these lines?

Carlos Quiles

The maps show simplified expansions of cultures depending on the most interesting events for each age. In this case, Vikings are over-represented for clarity. See here.

Tarno

Please pay close attention to Haplogroup N-PH1612.
Subgroup N-A9416 was tested and discovered to be closely related to the Ormenykut Conqueror, an elite burial.
Subgroup N-PH1896 was also tested and found to be closely related to the Tuzser Conqueror, also an elite burial.
Both sub-groups have deep Yakut ancestry and more about this will likely be published this calendar year. This will get very interesting.

Carlos Quiles

Cool. That’s from the most clearly Ugric-related N1c subclade, that didn’t appear in this paper, right?

Do you have a source?

Tarno

Tibor Feher will release a detailed paper this year on this. I have been provided sed research in advance. The Ormenykut and Tuzser are confirmed and will be published

The conqueror subclade depiction https://www.yfull.com/tree/N-PH1612/

Vinitharya

Hey, I’ve just read this paper by Csaba Horvath, a Hungarian scientist of Corvinus University and it is the first place, other than here, of the Uralic R1a link, and it makes the same conclusion of M458 lines spreading Slavic languages northeast into the Volga area and leading to the formation of Russia. I expect the Hungarians would eagerly adopt this, as it would mean their people are indigenous to Eastern Europe and not Trans-siberian nomads (believe me, I have seen a lot of Hungarians=Mongols memes)

Carlos Quiles

…this paper by Csaba Horvath… Yes, I reference him in the first paragraph of the Introduction of A Game of Clans. I expect the Hungarians would eagerly adopt this, as it would mean their people are indigenous to Eastern Europe and not Trans-siberian nomads I am not an expert in Hungarian nationalism, but I am guessing their old pan-Turanism and any other nativist theory they might have is not compatible with reality. This paper by professional geneticists is the best example, with their linking of Hungarians to Huns and Attila without any proof whatsoever…Or the previous one on their link… Read more »

Observer

The very daring conlusions of this study were criticied in Hungary as well by many scientists. However, we can not forget that the link between Huns and Hungarians is not only a romaticizied fairy tale invented recently, there are scientific facts (from the area of antropology, history etc), that can verify some links: We can use ancient Hungarian traditions written down in various sources. This tradition says: The first Hungarian rulers/kings (Árpád dynasty in 8??-1301) were direct descendants of Attila. The Hungarian tradition treats Attila and the Huns as positive heroes (in contradiction to the Western European tradition). An ancient… Read more »

[…] which point (together with later samples, such as those from modern Finno-Ugric peoples and Hungarian Conquerors) to the presence of basal subclades in the Andronovo-like Horizon, stemming from Abashevo migrants […]