On the origin and spread of haplogroup R1a-Z645 from eastern Europe

indo-european-uralic-migrations-corded-ware

In my recent post about the origin and expansion of haplogroup R1b-L51, Chetan made an interesting comment on the origin and expansion of R1a-Z645. Since this haplogroup is also relevant for European history and dialectal North-West Indo-European and Indo-Iranian expansion, I feel compelled to do a similar post, although the picture right now is more blurry than that of R1b-L51.

I find it interesting that many geneticists would question the simplistic approach to the Out of Africa model as it is often enunciated, but they would at the same time consider the current simplistic model of Yamna expansion essentially right; a model – if anyone is lost here – based on proportions of the so-called Yamnaya™ ancestral component, as found in a small number of samples, from four or five Eneolithic–Chalcolithic cultures spanning more than a thousand years.

The “75% Yamnaya ancestry of Corded Ware”, which has been given so much publicity since 2015, made geneticists propose a “Yamna → Corded Ware → Únětice / Bell Beaker” migration model, in order of decreasing Yamnaya proportions. Y-DNA and solid archaeological models suggested that this model was wrong, and recent findings have proven it was. In fact, the CWC sample closest to Yamna was a late outlier of Esperstedt in Central Europe, whose ancestry is most likely directly related to Yamna settlers from Hungary.

These wrong interpretations have been now substituted by data from two new early samples from the Baltic, which cluster closely to Yamna, and which – based on the Y-DNA and PCA cluster formed by all Corded Ware samples – are likely the product of female exogamy with Yamna peoples from the neighbouring North Pontic region (as we are seeing, e.g. in the recent Nikitin et al. 2018).

NOTE. There is also another paper from Nikitin et al. (2017), with more ancient mtDNA, “Subdivisions of haplogroups U and C encompass mitochondrial DNA lineages of Eneolithic-Early Bronze Age Kurgan populations of western North Pontic steppe”. Link to paper (behind paywall). Most interesting data is summarized in the following table:

yamna-corded-ware-mtdna

Even after the publication of Olalde et al. (2018) and Wang et al. (2018) – where expanding Yamna settlers and Bell Beakers are clearly seen highly admixed within a few generations, and are found spread across a wide Eurasian cline (sharing one common invariable trait, the paternally inherited haplogroup, as supported by David Reich) – fine-scale studies of population structure and social dynamics is still not a thing for many, even though it receives more and more advocates among geneticists (e.g. Lazaridis, or Veeramah).

NOTE. I have tried to explain, more than once, that the nature and origin of the so-called “Yamnaya ancestry” (then “steppe ancestry”, and now subdivided further as Steppe_EMBA and Steppe_MLBA) is not known with precision before Yamna samples of ca. 3000 BC, and especially that it is not necessarily a marker of Indo-European speakers. Why some people are adamant that steppe ancestry and thus R1a must be Indo-European is mostly related to a combination of grandaddy’s haplogroup, the own modern ethnolinguistic attribution, and an aversion to sharing grandpa with other peoples and cultures.

In the meantime, we are seeing the “Yamnaya proportion” question often reversed: “how do we make Corded Ware stem from Yamna, now that we believed it?”. This is a funny circular reasoning, akin to the one used by proponents of the Franco-Cantabrian origin of R1b, when they look now at EEF proportions in Iberian R1b-L23 samples. It seems too comic to be true.

R1a and steppe ancestry

The most likely origin of haplogroup R1a-Z645 is to be found in eastern Europe. Samples published in the last year support this region as a sort of cradle of R1a expansions:

  • I1819, Y-DNA R1a1-M459, mtDNA U5b2, Ukraine Mesolithic ca. 8825-8561 calBCE, from Vasilievka.
  • I5876, Y-DNA R1a, mtDNA U5a2a, Ukraine Mesolithic 7040-6703 calBCE, from Dereivka.
  • I0061, hg R1a1-M459 (xR1a1a-M17), mtDNA C1, ca. 6773-6000 calBCE (with variable dates), from Yuzhnyy Oleni Ostrov in Karelia.
  • Samples LOK_1980.006 and LOK_1981.024.01, of hg MR1a1a-M17, mtDNA F, Baikalic cultures, dated ca. 5500-5000 BC.
  • Sample I0433, hg R1a1-M459(xM198), mtDNA U5a1i, from Samara Eneolithic, ca. 5200-4000 BCE
  • Samples A3, A8, A9, of hg R1a1-M459, mtDNA H, from sub-Neolithic cultures (Comb Ware and Zhizhitskaya) at Serteyea, although dates (ca. 5th-3rd millennium BC) need possibly a revision (from Chekunova 2014).

NOTE. The fact that Europe is better sampled than North Asia, coupled with the finding of R1a-M17 in Baikalic cultures, poses some problems as to the precise origin of this haplogroup and its subclades. While the first (Palaeolithic or Mesolithic) expansion was almost certainly from Northern Eurasia to the west – due to the Mal’ta sample – , it is still unknown if the different subclades of R1a in Europe are the result of local developments, or rather different east—west migrations through North Eurasia.

Y-Full average estimates pointed to R1a-M417 formation ca. 6500 BC, TMRCA ca. 3500 BC, and R1a-Z645 formation ca. 3300 BC, TMRCA ca. 2900 BC, so the most likely explanation was that R1a-Z645 and its subclades – similar to R1b-L23 subclades, but slightly later) expanded quickly with the expansion of Corded Ware groups.

The presence of steppe ancestry in Ukraine Eneolithic sample I6561, of haplogroup R1a-M417, from Alexandria, dated ca. 4045-3974 calBCE, pointed to the forest steppe area and late Sredni Stog as the most likely territory from where the haplogroup related to the Corded Ware culture expanded.

However, the more recent Y-SNP call showing R1a-Z93 (L657) subclade rendered Y-Full’s (at least formation) estimates too young, so we have to rethink the actual origin of both subclades, R1a-Z93 (formation ca. 2900 BC, TMRCA ca 2700 BC), and R1a-Z283 (formation ca. 2900 BC, TMRCA ca. 2800 BC).

Contrary to what we thought before this, then, it is possible that the expansion of Khvalynsk-Novodanilovka chieftains through the steppes, around the mid-5th millennium BC, had something to do with the expansion of R1a-Z645 to the north, in the forest steppe.

We could think that the finding of Z93 in Alexandria after the expansion of Khvalynsk-Novodanilovka chiefs would make it more likely that R1a-Z645 will be found in the North Pontic area. However, given that Lower Mikhailovka and Kvitjana seem to follow a steppe-related cultural tradition, different to forest steppe cultures (like Dereivka and Alexandria), and that forest steppe cultures show connections to neighbouring northern and western forest regions, the rest of the expanding R1a-Z645 community may not be related directly to the steppe at all.

Adding a hypothetical split and expansion of Z645 subclades to the mid-/late-5th millennium could place the expansion of this haplogroup to the north and west, pushed by expanding Middle PIE-speaking steppe peoples from the east:

distribution-horse-scepters
Schematic depiction of the spread of horse-head scepters in the Middle Eneolithic, representing expanding Khvalynsk-Novodanilovka chieftains. See a full version with notes here.

The Złota culture

I have already written about the Podolia-Volhynia region: about the North Pontic steppe cultures in contact with this area, and about the chaotic period of migrations when Corded Ware seem to have first emerged there among multi-directional and multi-ethnic migrants.

This is what Włodarczak (2017) says about the emergence of Corded Ware with ‘steppe features’ after the previous expansion of such features in Central Europe with Globular Amphorae peoples. He refers here to the Złota culture (appearing ca. 2900-2800 BC) in Lesser Poland, believed to be the (or a) transitional stage between GAC and Corded Ware, before the emergence of the full-fledged “Corded Ware package”.

So far, to the north of the Carpathian Mountains, including Polish lands, no graves indicating their relationship with communities of the steppe zone have been found. On the contrary, the funeral rites always display a local, central European nature. However, individual elements typical of steppe communities do appear, such as the “frog-like” arrangement of the body (Fig. 20), or items associated with Pit Grave milieux (cf. Klochko, Kośko 2009; Włodarczak 2014). A spectacular example of the latter is the pointed-base vessel of Pit Grave culture found at the cemetery in Święte, site 11 near Jarosław (Kośko et al. 2012). These finds constitute a confirmation of the importance of the relationships between communities of Pit Grave culture and Corded Ware culture. They are chronologically diverse, although most of them are dated to 2600-2400 BC – that is, to the “classic” period of Corded Ware culture.

funnelbeaker-trypillia-corded-ware
Map of territorial ranges of Funnel Beaker Culture (and its settlement concentrations in Lesser Poland), local Trypillian groups and early Corded Ware Culture settlements (◼) at the turn of the 4th/3rd millennia BC.

However, when discussing the relationships with the steppe communities, Polish lands deserve particular attention since part of the groups inhabiting it belonged to the eastern province of Corded Ware culture (cf. Häusler 2014), which neighboured Pit Grave culture both from the east and south. In addition, there was a tradition of varied relationships with the north Pontic zone, which began to intensify from the second half of the 4th millennium BC (Kośko, Szmyt, 2009; Kośko, Klochko, 2009). These connections are especially readable in Małopolska and Kujawy (Kośko 2014; Włodarczak 2014). The emergence of the community of Globular Amphora culture in the north Pontic zone at the end of the 4th and the beginnings of the 3rd millennium BC (Szmyt 1999) became a harbinger of a cultural closening between the worlds of central Europe and the steppe.

The second important factor taking place at that time was the expansion of the people of Pit Grave culture in a westerly direction, along the Danube thoroughfare. As a result of this, also to the south of the Carpathian Mountains, e.g., along the upper Tisza River, a new “kurgan” cultural system was formed. As one outcome, the areas of central Europe, above all Małopolska, found themselves in the vicinity of areas inhabited by communities characterized by new principles of social organization and a new funeral rite. Around 2800 BC these changes became evident in different regions of Poland, with the most numerous examples being documented in south-eastern Poland and Kujawy. The nature of the funeral rite and the features of the material culture perceptible at that time do not have straight forward analogies in the world of north Pontic communities. In this respect, the “A-horizon” is a phenomenon of local, central European origin. The events preceding the emergence of the said horizon (that is, the expansion of the people of Pit Grave culture into the area north of the arc of the Carpathians) are nowadays completely unidentifiable and remain merely an interesting theoretical matter (cf. e.g., Kośko 2000). Therefore, analysis of the archaeological sources cannot confirm the first archaeogenetic analysis suggesting a bond between the communities of the Pit Grave culture and Corded Ware culture (e.g., Haak et al. 2015).

Artefacts of the “A-horizon”, i.e., shaft-hole axes, amphorae (Fig. 21), beakers, and pots with a plastic wavy strip (Fig. 7) are found in different funerary and settlement contexts, sometimes jointly with finds having characteristics of various cultures (e.g., in graves of Złota culture, or at settlements of Rzucewo culture). Hence, they primarily represent a chronological phase (c. 2800-2600 BC), one obviously related to the expansion of a new ideology.

Eastern CWC expansion

Before continuing tracing the Corded Ware culture’s main features, it is worth it to trace first their movement forward in time, as Corded Ware settlers, from Poland to the east.

Circum-Baltic CWC

According to Klochko and Kośko (1998):

The colonizing Neolithic waves are continued by the Circum-Baltic Corded Ware culture, closely related to the traditions of the Single Grave culture and traditions of the Northern European Lowlands. After ca. 2900 BC, certain cultural systems with ‘corded’ traits –genetically related to the catchment area of the south-western Baltic – appear in the drainages of the Nemen, Dvina, Upper Dnieper, and even the Volga. These communities are considered the vector of Neolithisation in the Forest Zone.

east-european-fatyanovocwc
East European movement directions (arrows) of the representatives of the Central European Corded Ware Culture. Modified from I.I. Artemenko.

The picture in the Baltic (Pamariu / Rzucewo) and Finland (Battle Axe) is thus more or less clearly connected with early dates ca. 2900-2800 BC:

There is a clear interaction sphere between the eastern Gulf of Finland area – reaching from Estonia to the areas of present-day Finland and the Karelian Isthmus in Russia –, evidenced e.g. by the sharp-butted axes, derived from the Estonian Karlova axe.

Interesting in this regard is the expansion of the Corded Ware culture in Finland, into a far greater territory than previously thought, that is poorly represented in most maps depicting the extent of the culture in Europe. Here is summary of CWC findings in Finland, using images from Nordqvist and Häkäla (2014):

finland-cwc
Corded Ware culture remains in Finland, excluding the so-called ‘imitations’. [Notice in the top left image the often depicted border of the culture]. Combination of maps from Nordqvist & Häkälä (2014)

Middle Dnieper and Fatyanovo

The earliest Middle Dnieper remains are related to CWC graves between the Upper Vistula and the Bug, containing pottery with Middle Dnieper traits, dated probably ca. 2700 BC, which links it with the expansion of the A-horizon. In fact, during the period ca. 2800-2400 BC, the area of Lesser Poland (with its numerous kurgans and catacomb burials) is considered the western fringe of an area spreading to the east, to the middle Dniester and middle Dnieper river basins, i.e. regions bordering the steppe oecumene. This ‘eastern connection’ of funeral ritual, raw materials, and stylistic traits of artefacts is also identified in some graves of the Polish Lowlands (Włodarczak 2017).

gac-cwc-baltic-dnieper
Cultural situation in Eastern Europe in approximately the middle of the III mill. BC. Key: 1 – areas settled by Globular Amphora culture populations; 2 – areas penetrated by Globular Amphora culture populations; 3 – border between central and eastern group; 4 – Pamariu/Rzucewo culture area; 5 – zone of Pamariu/Rzucewo culture influences; 6 – directions of Comb Pottery culture influence; 7 – Zhizhitskaya culture; 8 – eastern border of “pure” Corded Ware site; 9 – North Belarussian culture; 10 – Middle Dnieper culture; 11 – Fatyanovo culture; 12 – Yamnaya culture; 13 – eastern border of Dniester group; 14 – Kemi-Oba culture and influences; 15 – Foltesti culture; 16 – syncretic sites with evidence of Globular Amphora culture traits (1 – Nida; 2 – Butinge; 3 – Palanga; 4 – Juodkrante; 5 – Azyarnoye; 6 – Mali Rogi; 7 – Prorva; 8 – Strumen/Losha; 9 – Syabrovichi; 10 – Luchin-Zavale; 11 – Lunevo (?); 12 – Belynets; 13 – Losiatyn; 14 – Corpaci; 15 – Ocnita; 16-17 – Camenca; 18 – Marculesti; 19 – Orhei; 20 – Efimovka; 21 – Tatarbunary; 22 – Novoselitsa; 23 – Primorskoye; 24 – Sanzhiyka; 25 – Akkermen; 26 – Maydanetskoye; 27 – Grigorevka; 28 – Kholmskoye; 29 – Purcari; 30 – Roscani; 31 – Semenovka; 32 – Grishevka; 33 – Durna Skela; 34 – Iskovshchina; 35 – Primorskoye); 17 – borders of ecological zones. From Szmyt (2010)

The Fatyanovo (or Fatyanovo-Balanovo) culture was the easternmost group of the Corded Ware culture, and occupied the centre of the Russian Plain, from Lake Ilmen and the Upper Dnieper drainage to the Wiatka River and the middle course of the Volga. From the few available dates, the oldest ones from the plains of the Moskva river, and from the late Volosovo culture containing also Fatyanovo materials, and in combination they show a date of ca. 2700 BC for its appearance in the region. The Volosovo culture of foragers eventually disappeared when the Fatyanovo culture expanded into the Upper and Middle Volga basin.

The origin of the culture is complicated, because it involves at its earliest stage different Corded Ware influences in neighbouring sites, at least on the Moskva river plains (Krenke et al. 2013): some materials (possibly earlier) show Circum-Baltic and Polish features; other sites show a connection to western materials, in turn a bridge to the Middle Dnieper culture. This suggests that groups belonging to different groups of the corded ware tradition penetrated the Moscow region.

The split of subclades Z93 – Z283

If we take into account that the split between R1-Z93 and R1a-Z283 must have happened during the 5th millennium BC, we have R1a-Z93 likely around the middle Dnieper area (as supported by the Alexandria sample), and R1a-Z283 possibly to the north(-west), so that it could have expanded easily into Central Europe, and – through the northern, Baltic region – to the east.

Where exactly lies the division is unclear, but for the moment all reported Circum-Baltic samples with Z645 subclades seem to belong to Z282, while R1a samples from Sintashta/Potapovka (including the Poltavka outlier) point to Abashevo being dominated by R1a-Z93 subclades.

We have to assume, then, that an original east-west split betwen R1a-Z283 and R1a-Z93 turned, in the eastern migrations, into a north-south split between Z282 and Z93, where Finland and Battle Axe in general is going to show Z282, and Middle Dnieper – Abashevo Z93 subclades.

copper-age-early-cwc
Early Copper Age migrations ca. 3100-2600 BC.

I can think of two reasons why this is important:

  1. Depending on how Proto-Corded Ware peoples expanded, we may be talking about one community overcoming the other and imposing its language. Because either
    • clans of both Z93 and Z283 were quite close and kept intense cultural contacts around Dnieper-Dniester area; or
    • if the split is as early as the 5th millennium BC, and both communities separated then without contact, we are probably going to see a difference in the language spoken by both of them.
  2. In any case, the main north-south division of eastern Corded Ware groups is pointing to an important linguistic division within the Uralic-speaking communities, specifically between a Pre-Finno-Ugric and a Pre-Samoyedic one, and potentially between Pre-Finno-Permic and Pre-Ugric.

These may seem irrelevant questions – especially for people interested only in Indo-European migrations. However, for those interested in the history of Eurasian peoples and languages as a whole, they are relevant: even those who support an ‘eastern’ origin of Proto-Uralic, like Häkkinen, or Parpola (who are, by the way, in the minority, because most Uralicists would point to eastern Europe well before the Yamna expansion), place the Finno-Ugric expansion with the Netted Ware culture as the latest possible Finno-Ugric immigrants in Fennoscandia.

The Netted Ware culture

The image below shows the approximate expansion of Corded Ware peoples of Battle Axe traditions in Finland, as well as neighbouring Fennoscandian territories, from ca. 2800 BC until the end of the 3rd millennium. A controversial 2nd (late) wave of the so-called Estonian Corded Ware is popular in texts about this region, but has not been substantiated, and it seems to be a regional development, rather than the product of migrations.

finland-corded-ware
Left: Corded Ware remains in Finland from ca. 2800 BC, according to Nordqvist & Häkälä (2014), combined in a single image. Right: Distribution of the Corded Ware culture within Finland. Mapped (black dots) are finds of typical stone battle axes, used as a proxy (data from [8]). The red isolines indicate average permanent snow cover period from 1981 to 2010 (data from [9]). A recent study estimates the snow cover period ca 4500 years ago would have been 40–50 days less than today [10]. Overlying coloration refers to the lactose persistance (LP) allele gradient in modern northeastern Europe (see the electronic supplementary material, appendix B: Material and methods and table 1, for details); lozenge dots specify the dataset mean points for the triangulation. From Cramp et al. (2014).

As we have seen, Fatyanovo represents the most likely cultural border zone between Circum-Baltic peoples reaching from the Russian Battle Axe to the south, and Middle Dnieper peoples reaching from Abashevo to the north. In that sense, it also represents the most likely border culture between north-western (mainly R1a-Z282) and south-eastern (R1a-Z93) subclades.

With worsening climatic conditions (cooler seasons) at the end of the 3rd millennium, less settlements are apparent in the archaeological record in Finland. After ca. 2000 BC, two CWC-related cultures remain: in the coast, the Kiukainen culture, derived from the original Circum-Baltic Corded Ware settlers, reverts to a subsistence economy which includes hunting and fishing, and keeps mainly settlements (from the best territories) along the coast. In the inland, Netted Ware immigrants eventually appear from the south.

cultures-western-finland
Image modified from Cramp et al. (2014) “The timeline shows the archaeological cultures
discussed here alongside actual sherds sampled and typical vessel forms (after [26–28]) (latter not shown to scale). Distribution maps show the geographical range of (f) Typical Comb Ware, (g) Corded Ware, (h) Kiukainen Ware and (i) Bronze Age cultures in the region (after [10,20,29]).”

The Netted Ware culture emerged in the Upper Volga–Oka region, derived from the Abashevo culture and its interaction with the Seima-Turbino network, and spread ca. 1900-1800 BC to the north into Finland, spreading into eastern regions previously occupied by cultures producing asbestos and organic-tempered wares (Parpola 2018).

NOTE. Those ‘contaminated’ by the Copenhagen fantasy map series may think that Volosovo hunter-gatherers somehow survived the expansion of Fatyanovo-Balanovo and Abashevo, hidden for hundreds of years in the forest, and then reappeared and expanded the Netted Ware culture. Well, they didn’t. At least not in archaeological terms, and certainly not with the genetic data we have.

If we combine all this information, and we think about these peoples in terms of Pre-Finno-Permic and Pre-Ugric languages developing side by side, we get a really interesting picture (see here for Proto-Fennic estimates):

  • The Battle Axe around the Baltic Sea – including the Gulf of Finland and Scandinavia – would be the area of expansion of Pre-Finno-Permic peoples, of R1a-Z283 subclades, which became later concentrated mainly on coastal regions;
  • the southern areas may correspond to Pre-Ugric peoples, which expanded later to the north with Netted Ware (see image below) – their precise subclades may be dependent on what will be found in Fatyanovo;
  • and Pre-Samoyedic peoples (of R1a-Z93 subclades) would have become isolated somewhere in the Cis- or (more likely) Trans-Urals region after 2000 BC, possibly from the interaction of the latest Balanovo stages and the Seima-Turbino phenomenon.
netted-ware-parpola
Distribution of the Netted Ware according to Carpelan (2002: 198). A: Emergence of the Netted Ware on the Upper Volga c. 1900 calBC. B: Spread of Netted Ware by c. 1800 calBC. C: Early Iron Age spread of Netted Ware. (After Carpelan 2002: 198 > Parpola 2012a: 151.)

These communities in contact would have allowed for:

  • the known Indo-Iranian loanwords in Finno-Ugric to spread through a continuum of early dialects formed by Abashevo – Fatyanovo – Battle Axe groups;
  • the Finno-Saamic substrate of Germanic to be associated with Battle Axe groups in Scandinavia;
  • the important Palaeo-Germanic loanwords in Finno-Saamic spreading with long-term contacts (from Pre-Germanic to the Proto-Germanic, and later North Germanic period) through the Baltic Sea, between Scandinavia and the Gulf of Finland;
  • and Tocharian contacts with Samoyedic (although limited, and in part controversial), which point to its early expansion to the east of the Ural Mountains.

On the other hand, if one is inclined to believe that R1a and steppe ancestry do represent Indo-European speakers… which language was spoken from the Gulf of Finland well into the north, the inland, and Karelia, and in Northern Russia, by Corded Ware peoples and their cultural heirs (like Kiukainen or Netted Ware) for almost three thousand years?

Because we know that no other peoples of different haplogroups dominated over eastern Fennoscandia until the Iron Age, and N1c and Siberian ancestry expanded separately, and probably due to late bottlenecks, especially with Fennic peoples expanding recently to the north at the expense of the Saami population.

After the expansion of Bell Beaker peoples, the geographic distribution of late Corded Ware groups in the second half of the 3rd millennium, just before their demise – and before the expansion of Netted Ware to the north – , can be depicted thus as follows:

bronze_age_early_cut
Early Bronze Age Europe.

Territories in cyan must then represent, for some people who believe in an archaic Indo-Slavonic of sorts, the famous Fennoscandian Balto-Slavic to the north (before they were displaced by incoming Finno-Saamic peoples of hg N1c during the Iron Age and up to the Middle Ages); and the also famous Tundra-Forest Indo-Iranian in the Upper Volga area, a great environment for the development of the two-wheeled chariot…

But let’s leave the discussion on imaginary IE dialects for another post, and continue with the real question at hand.

A steppe funerary connection?

Back to Złota as a transitional culture, we have already seen how the corded ware vessels characteristic of the Classic CWC are related to Globular Amphora tradition, and show no break with this culture. It is usually believed that the funerary rites were adopted from steppe influence, too. That is probably right; but it does not mean that it came from Yamna or other coeval (or previous) steppe culture; at least not directly.

NOTE. A similar problem is seen when we read that Mierzanowice or Trzciniec show “Corded Ware” traits from a neighbouring CWC group, when CWC groups disappeared long before these cultures emerged. For cultural groups that are separated centuries from each other, an assertion as to their relationship needs specifics in terms of dates and material connection, or it is plainly wrong.

These are the funerary ritual features from Złota (later specialized in Corded Ware), as described by Włodarczak (2017):

  • Single burial graves; along with the habit of interring the deceased in multiple burial graves, but emphasizing their individual character by careful deposition of the body and personal nature of the grave goods.
  • Grave goods with materials and stylistiscs belonging to an older system (e.g. amber products); and others correlated to the ‘new world’ of the CWC, such as flint products made of the raw materials tipical of Lesser Poland’s CWC, copper ornaments, stone shaft-hole axes, bone and shell ornaments, and characteristic forms of vessels like beakers and amphoras.
  • Military goods, which would become prevalent in later periods, are present in a moderate number, compatible with their lesser importance.
  • There are also cases of the characteristic catacomb (“niche”) graves – with an entrance pit, a more extensive niche, and a narrow corridor leading to a vault – , as well as some individual cases of application of ochre and deformation of skulls.
catacomb-grave-ksiaznice
Catacomb grave no. 2a/06 from Książnice, Złota culture (acc. to Wilk 2013). Image from Włodarczak (2017)

It seems that the Złota funerary tradition was also “transitional”, like corded ware vessels, into the classical Corded Ware ideology. But “transitional” from what exactly? Yamna? Probably not.

The Lublin-Volhynia culture

One needs not look for a too distant culture to find similarities. Włodarczak (2017) points to CWC in south-eastern Poland and Kuyavia showing, by the time of the Yamna expansion, a funeral rite and features of the material culture without straightforward analogies in the world of north Pontic communities, and thus suggests that the “A-horizon” is a local phenomenon of central European origin.

This assertion is interesting, in so far as most Corded Ware samples investigated to date seem to come precisely from an East-Central territory near the Ukraine forest steppe, with a cluster already established by the end of the 5th millennium:

PCA-caucasus-lola-ane-chg
Image modified from Wang et al. (2018). Samples projected in PCA of 84 modern-day West Eurasian populations (open symbols). Previously known clusters have been marked and referenced. An EHG and a Caucasus ‘clouds’ have been drawn, leaving Pontic-Caspian steppe and derived groups between them.See the original file here.

The following text is from Stanisław Wilk (2018), about the Lublin-Volhynian (and related) cemeteries at Wyciąże and Książnice:

lublin-volhynia-culture
A reach of the Wyciąże-Złotniki group and Lublin-Volhynian culture in the south-eastern Poland and western Ukraine: 1. Area of the Wyciąże-Złotniki group; 2. Area of the Lublin-Volhynian culture. A. Cemetery of the Lublin-Volhynian culture at site 2 in Książnice; B. Cemetery of the Wyciąże-Złotniki group at site 5 in Kraków Nowa Huta-Wyciąże (drawing by S. Wilk based on Zakościelna 2006 and Nowak 2014, on a background downloaded from https://maps-for-free.com/).

Regardless of the differences between the two necropolises (such as the number of burials, the area which has been explored, the orientation and layout of burials), it seems that they have several key elements in common:

  • concentration of graves in separate cemeteries;
  • differentiation of burials with regard to sex (the principle of the ‘left ̶ right’ side, different burial goods for males and females);
  • stratification of graves with regard to the richness of their inventories (this mainly applied to copper artefacts);
  • occurrence of indicators of the richest male burials (a copper dagger in Wyciąże, a copper battle axe, a small axe and a chisel in Książnice);
  • allocation of a separate area for elite burials (the eastern burial area in Książnice, and the southeastern and north-central part of the necropolis in Wyciąże), as well as one for egalitarian burials (the western area in Książnice, and the south-central and western part of the cemetery in Wyciąże).
lublin-volhynian-eneolithic-cemetery
Plan of the Lublin-Volhynian culture cemetery at site 2 in Książnice: 1. female graves; 2. man graves; 3. copper traces; 4. cenothap; 5. cremation grave; 6. partial grave; 7. estimated area of the L-VC cemetery; 8. estimated area of an elite and poor burial fields; 9. area of burials containing copper artefacts (drawing by S. Wilk).

The above-mentioned characteristics prove that the patterns of social and religious behaviours from areas lying beyond the Carpathian Mountains exerted a strong influence on the two societies living in Lesser Poland.

Anna Zakościelna, while describing the similarities between the burial ritual of the late Polgár groups and cultures from areas on the Tisza river and the Lublin-Volhynia culture, claimed that:

a characteristic feature of the burial ritual of both cultures was practicing various group norms, which required different treatment of the deceased depending on their sex, age and social rank. As in the Lublin-Volhynia culture, the opposition ‘male – female’ can the most clearly be observed ̶ particularly, in the consistent positioning of males on the right, and females, on the left side. And, there is much indication that this ritual norm divided the deceased from early childhood (Sofaer Derevensky 1997: 877, Tab. 1; Lichter 2001: 276- 280, 322-323) (Zakościelna 2010: 227-228).

It seems that these observations can also be extended to the Wyciąże-Złotniki group.

Another question is whether the evidence of the influences of the copper civilization observed in both cemeteries emerged as a result of the literal copying of patterns from the south, or whether the latter were only a source of inspiration for local solutions.

Looking at this problem form the perspective of the details of burial ritual, between the Carpathian Basin and Lesser Poland, we can observe clear differences, among others, in the size of cemeteries and orientation of burials. While, in the Carpathian Basin there were large necropolises, consisting of several dozen burials located in rows, with the dominant orientation along the SE-NW and E-W axis (Lichter 2001: Abb. 123, 143; Kadrow 2008: 87); in Lesser Poland there were small cemeteries of several to a dozen or so burials, mostly oriented along the S-N axis (in the Lublin-Volhynia culture; Zakościelna 2010: 66), as well as S-E and NE-SW (in the Wyciąże-Złotniki group; Kaczanowska 2009: 77). Similarly, there are differences in the details of the burial goods. North of the Carpathians, there is a much smaller frequency of copper artefacts, particularly in the group of prestigious, heavy items (battle axes, axes and daggers), as well as a complete lack of objects made of gold. Want is more, the pottery found in the graves has a distinct local character, only supplemented by imitating or imports from areas beyond the Carpathians (Zakościelna 2006: 85; Nowak 2014: 273; a different opinion Kozłowski 2006: 57). Therefore, the suggestion made by Nowak seems right ̶ namely, that these influences were not caused by migrations of groups of the population living on the Tisza river to Lesser Poland, but were rather due to processes of selective cultural transmission (Nowak 2014: 273).

Therefore, the sharing of a similar funerary rite (as happened later between Lublin-Volhynia and Złota), although it shows a strong cultural connection with autochthonous cultures, is obviously not the same as sharing ancestors; and even if it were so, they would not need to be paternal ancestors. But it shows that important Corded Ware cultural traits are local developments, and it disconnects thus still more supposed CWC ‘steppe traits’ from steppe cultures, and connects them with the first steppe-related cultural wave that reached central Europe in the 5th millennium BC.

Prehistoric Pontic—Caspian links

How would a Lublin-Volhynia culture be related to the North Pontic area ca. 4500-3000 BC? We can enjoy the map series of Baltic—Pontic migrations by Viktor Klochko (2009), and make a wild guess:

baltic-pontic-routes
Pontic—Baltic routes of migrations during the Eneolithic. Top left: Linear Pottery expansion. Top right: Funnel Beaker expansion. Bottom left: late Trypillia expansion. Bottom right: GAC expansion.

And then read the account of Sławomir Kadrow, in Exchange of People, Ideas and Things between Cucuteni-Trypillian Complex and Areas of South-Eastern Poland (2016):

In the second half of the 5th millennium BC (horizon 1), communities of the Tripolye culture, phases BI-BII, had contacts with the population of the late (IIa) phase of the Malice culture. The areas settled by both cultural complexes were located at a great distance from each other. The communities of the Tripolye culture adopted selected features of Malice ceramic production (fig 2). This seems to have resulted from marital exchange: on a moderate scale, Tripolye men sought out their wives in the area of the Malice culture and, according to patrilocal marriage customs, the women then moved to the Tripolye settlements, sporadically transferring ready-made ceramic products, so-called imports, to the Tripolye culture. Thus, the wives were responsible for the considerably more numerous imitations of the Malice ceramics and the long-lasting, though selective, traditions of Malice pottery passed down in their new environment. The patrilocal marriage customs involving the Malice women and the Tripolye men (never the other way round), and the fact that pottery was women’s domain, led to the unidirectional transfer of vessels, technology and norms of ceramic production from the Malice culture to the Tripolye culture.

The turn of the 5th and the 4th millennia and the early 4th millennium BC (horizon 2) witnessed the deepening interaction between the populations of the youngest (IIb) phase of the Malice culture and the classic (II) phase of the Lublin-Volhynia culture on the one hand and the communities of phase BII of the Tripolye culture on the other. The Danube and the Tripolye settlement complexes came into contact on the upper Dniester and between the Styr and the Horyn rivers in Volhynia. This helped to continue the previous forms of marital exchange, which resulted in further popularisation of the ceramics and the traditions of ceramic production typical of the Danube cultures, i.e. the Malice and the Lublin-Volhynia cultures, and also the Polgár culture, in the areas settled by the Tripolye cultural complex.

As the civilizational norms of the Eneolithic (Copper) Age became widespread in that period, the forms of interaction described above acquired new elements. The deepening internal diversification of the early Eneolithic communities of the Lublin-Volhynia culture led to a growing demand for prestige objects, which was met with import or imitation of copper artefacts, mainly those from the Carpathian Basin, and with flint tools produced from long blades. That type of flint production depended largely on new technologies derived from the Tripolye culture, as proven by such borrowings as troughlike retouch or the very idea and technology for the production of long flint blades in the Lublin-Volhynia culture. It seems that the influx of Tripolye settlers into flintbearing areas in Volhynia and on the upper Dniester, adjacent to the settlement centres of the late phase of the Malice culture and the Lublin-Volhynia culture, created sufficient conditions for the expanding influence of the Tripolye flint working on the communities of the Eneolithic Lublin-Volhynia culture.

In the mid-4th millennium BC (horizon 3), those forms of interaction between the Danube communities (the late phase of the Lublin-Volhynian culture) and the Tripolye communities (phase CI)were continued. Elements of the Danube pottery still grew in popularity in the Tripolye population, while selected features of the Tripolye flint working were adopted by the Lublin-Volhynia culture.

In that period, the population of the Funnel Beaker culture of the pre-classic and early classic phases (the beginnings of Gródek 1 and Bronocice III), until then absent from those areas, quite quickly drove out and replaced the Danube population in western Volhynia and the upper Dniester basin. This caused significant changes in the forms and intensity of the intercultural interaction, which became fully apparent already in the 2nd half of the 4th millennium BC.

In the following period (horizon 4), the population of the classic phase of the Funnel Beaker culture (Gródek 1, Bronocice III) settled more and more intensively the upper Dniester basin, up to the Hnyla Lypa river, and western Volhynia, up to the Styr river. East of those rivers, the Funnel Beaker settlers created considerable areas where they mixed with settlers from early phase CII of the Tripolye culture. Their coexistence, lasting there for many generations, resulted in deepening the interactions between members of both cultural complexes and in developing entirely new forms of relationships.

(…)

The intensifying interaction between the communities of the Funnel Beaker culture and the Tripolye culture, early phase CII, in the 2nd half of the 4th millennium BC (horizon 4) was an introduction to, and perhaps a condition for, even more frequent contacts in the next period, the first centuries of the 3rd millennium BC (horizon 5). In that case, the interaction was mainly triggered by multidirectional migrations of larger human groups, involving a significant part of the population of all cultures from the areas discussed here. The Tripolye communities of younger phase CII settled Volhynia, its eastern areas in particular, from the south and the south-east, while groups representing the younger phases of the Funnel Beaker culture (Gródek 2), often with Baden features (Bronocice IV and V), moved increasingly into the western part of that region. The Yamna communities expanded along the lower and central Danube to the west, whereas the populations of the late phase of the Baden culture took the opposite direction, reaching as far as Kiev in the northeast, and contributed to the cultural character of the Sofievka group.

The communities of the Globular Amphora culture migrated from the north-west, from eastern Poland, towards the Danube Delta and as far as the Dnieper in the east, while the multicultural population from the areas around the mouth of the Danube moved in the opposite direction, carrying with them cultural elements from Thrace, or even from Anatolia. Some of them returned to the starting point (to south-eastern Poland), bringing with them a new form of pottery, so-called Thuringian amphora, borrowed from the late Trypillian Usatovo group. This resulted in origins of the Złota culture, a cultural phenomenon that gave beginnings to the oldest Corded Ware culture. Inventories of both cultures contained the already mentioned Thuringian amphorae.

lublin-volhynia-alexandria
Graves and cemeteries with gender differentiated burial rites in Europe; A — Hamangia and Varna cultures; B — Tiszapolgar and Bodrogkeresztur cultures; C — Lublin-Volhynia culture; D — Brześć Kujawski culture. Added star symbol with approximate location of the Alexandria site. Modified image from Sławomir Kadrow (2016)

Here is a more recent assessment (2017) of the latest radiocarbon analyses of the available settlements of cultures in the area, published by Marek Novak (announced in a previous post), which gives the following data on Wyciąże-Złotniki, Lublin-Volhynia, and Wyciąże/Niedźwiedź:

This scheme unambiguously suggests both the overlapping and contiguous nature of cultural development in western Lesser Poland within the Middle Neolithic. The basic elements of this development are: 1) the Wyciąże-Złotniki group and the Lublin-Volhynian culture, until c. 3650–3550 cal BC; 2) the Funnel Beaker culture proper, which appeared c. 3750–3700c al BC, and existed until c. 3300–3250 cal BC, perhaps accompanied by the Wyciąże/Niedźwiedź materials from c. 3650–3550 cal BC; and 3) the Baden culture and the Funnel Beaker/Baden assemblages from 3100 and 3300–3100 cal BC, respectively, until 2850–2750 and 2850 cal BC, with – possibly – later Funnel Beaker culture and Wyciąże/ Niedźwiedź materials, existing until c. 3100 cal BC.

The final scheme shows that the Lublin-Volhynian culture could have coincided with the Wyciąże-Złotniki group. In view of the territorial relationship between them, relations from the point of view of material culture, primarily in the field of pottery, become particularly interesting. It is relatively easy to see clear similarities between these units. However, the most evident similarities apply only to some categories of ceramics, including, for example, vessels with Scheibenhenkel handles. What is more, in the period between the late 38th and early 36th centuries BC, the early Funnel Beaker and possibly early Baden influences are superimposed on this Lublin-Volhynian/Wyciąże-Złotniki ‘mix’.

[About Corded Ware: The] development of this unit in central Europe, including western Lesser Poland, [] usually point to c. 2800 cal BC (Włodarczak 2006a). (…) the calibration curve makes it possible to alternatively refer several dates earlier than c. 3100 to c. 2850–2800 cal BC.

Conclusion

There is no direct archaeological link of Lublin-Volhynia-related groups with Corded Ware, beyond the fact that they shared homeland and Central European (‘steppe-related’) traits, as found in the Złota culture. But there is no direct link of Yamna with Corded Ware, either, whether in terms of culture or population.

So, given the evident link of R1a-Z93 and steppe ancestry with the forest steppe ca. 4000 BC, the surrounding North Pontic areas in contact along the Dniester, Dnieper, Bug, and Prut are the best candidates for the appearance of R1a-Z283: steppe cultures to the south and south-west; sub-Neolithic (Comb Ware) groups to the north in the forest zone; and Eneolithic groups to the west and north-west.

Seeing how ‘ancestral components’ and PCA cluster can change within a few generations, the question of the spread of R1a-Z645 subclades is still not settled by a single sample in Alexandria. However, based on the explosive expansions we are seeing from small territories, it would not be surprising to find R1a-Z93 and R1a-Z283 side by side in the same small area within the forest steppe.

NOTE. An archaeological link may not mean anything relevant in genetics, especially – as in this case – when no clear migration event has been traced to date. We have seen exactly that with Kristiansen’s proposal of a long-term genetic admixture of Yamna with Trypillia and GAC to form Corded Ware, which didn’t happen. The cultural and ideological connection of CWC peoples with Lublin-Volhynian tradition may be similar to the already known connection with GAC, and not mean anything in genetic finds; at least in terms of Y-DNA haplogroup.

We believed in the 2000s that Corded Ware represented the expansion of Late Proto-Indo-European, because the modern map of haplogroup R1a showed a distribution similar to how we thought the European and Indo-Iranian languages could have expanded. This has been proven wrong, and that’s what ancient DNA is for; not to confirm the own ideas or models, or to support modern ideologies.

It is impossible to know if R1a-Z645 comes from the steppe, forest steppe, or forest zone, until more samples are published. I don’t think there will be any big surprise, no matter where it is eventually found. By now, adding linguistic reconstruction to archaeological traits, and to the genetic data from Yamna and Corded Ware settlers, the only clear pattern is that patrilineal clans expanded, during the Final Eneolithic / Chalcolithic:

  • Late Proto-Indo-European with Yamna and R1b-L23 subclades, given the known genomic data from Khvalynsk, Yamna, Afanasevo, Bell Beaker, Catacomb, and Poltavka—Sintashta/Potapovka.
  • Uralic with Corded Ware and R1a-Z645 subclades, given the known genomic data from Fennoscandia and the Forest Zone.

Everything else is just wishful thinking at this moment.

Related

The Indo-European demic diffusion model, and the “R1b – Indo-European” association

yamna_bell_beaker_cut

Beginning with the new year, I wanted to commit myself to some predictions, as I did last year, even though they constantly change with new data.

I recently read Proto-Indo-European homelands – ancient genetic clues at last?, by Edward Pegler, which is a good summary of the current state of the art in the Indo-European question for many geneticists – and thus a great example of how well Genetics can influence Indo-European studies, and how badly it can be used to interpret actual cultural events – although more time is necessary for some to realize it. Notice for example the distribution of ‘Yamnaya’ in 3000 BC, all the way to Latvia (based on the initial findings of Mathieson et al. 2017), and the map of 2000 BC with ‘Corded Ware’, both suggesting communities linked by admixture and unrelated to actual cultures.

Some people – especially those interested in keeping a simplistic picture of Europe, either divided into admixture groups or simplistic R1b-Vasconic / R1a-Indo-European / N1c-Uralic (or any combination thereof) – want (others) to believe that I am linking ‘Indo-Europeans’ with haplogroup R1b. That is simply not true. In fact, my model dismisses such simplistic identifications of the reconstructible proto-languages with any modern peoples, admixtures, or haplogroups.

vasconic-uralic
Simplistic Vasconic/R1b-Uralic/N1c distribution, and intruding Indo-European/R1a, according to Wiik.

The beauty of the model lies, therefore, precisely in that if you take any modern group speaking Indo-European languages, none can trace back their combination of language, admixture, and/or haplogroup to a common Indo-European-speaking people. All our ancestral lines have no doubt changed language families (and indeed cultures), they have admixed, and our European regions’ paternal lines have changed, so that any dreams of ‘purity’ or linguistic/cultural/regional continuity become absurd.

That conclusion, which should be obvious to all, has been denied for a long time in blogs and forums alike, and is behind the effort of many of those involved in amateur genetics.

Main linguistic aim

The main consequence of the model, as the title of the paper suggests, is that reconstructible Indo-European proto-languages expanded with people, i.e. with actual communities, which is what we can assert with the help of Genomics. From a personal (or ethnic, or political) point of view genomics is useless, but from an anthropological (and thus linguistic) point of view, genomics can be a very useful tool to decide between alternative models of language diffusion, which has given lots of headaches to those of us involved in Indo-European studies.

The demic diffusion theory for the three main stages of the proto-language expansion was originally, therefore, a dismissal of impossible-to-prove cultural diffusion models for the proto-language – e.g. the adoption of Late Proto-Indo-European by Corded Ware groups due to a patron-client relationship (as proposed by Anthony), or a long-lasting connection between cultures (as proposed by Kristiansen, and favoured by “constellation analogy” proponents like Clackson, who negated the existence of common proto-languages). It also means the acceptance of the easiest anthropological model for language change: migration and – consequently – replacement.

By the time of the famous 2015 papers, I had been dealing for some time with the idea that the shared features between Indo-Iranian and Balto-Slavic may have been due to a common substrate, and must have therefore had some reflection in genomic finds. The data on these papers, and the addition of a weak connection between Pre-Germanic and Balto-Slavic communities, together with their clearest genetic link – R1a-M417 subclades (especially European Z283) – made it still easier to propose a Corded Ware substrate, partially common to the three.

Allentoft Corded Ware
Allentoft et al. “Arrows indicate migrations — those from the Corded Ware reflect the evidence that people of this archaeological culture (or their relatives) were responsible for the spreading of Indo-European languages. All coloured boundaries are approximate.”

Before the famous 2015 papers (and even after them, if we followed their interpretation), we were left to wonder why the supposed vector of expansion of Indo-European languages, Corded Ware migrants – represented by R1a-Z645 subclades, and supposedly continued unchanged into modern populations in its ‘original’ ancestral territories, Balto-Slavic and Indo-Iranian – , were precisely the (phonetically) most divergent Indo-European languages – relative to the parent Late Indo-European proto-language.

My paper implied therefore the dismissal of an unlikely Indo-Slavonic group, as proposed by Kortlandt, and of a still less factible Germano-Slavonic, or Germano-Indo-Slavonic (?) group, as loosely implied by some in the past, and maybe supported in certain archaeological models (viz. Kristiansen or partially Anthony), and presently by some geneticists since their simplistic 2015 papers on “massive migrations from the steppe“, and amateur genetic fans with infinite pet theories, indeed.

A common Corded Ware substrate to Balto-Slavic and Indo-Iranian, and common also partially between Balto-Slavic and Germanic (as supported by Kortlandt, too, albeit with different linguistic connotations), would explain their common features. The Corded Ware culture (and Uralic, tentatively proposed by me as the group’s main language family) is a strong potential connection between them, further supported by phylogeography, too.

Other consequences

Interpretations in my paper help thus dismiss the simplistic Yamna -> Corded Ware -> Bell Beaker migration model implied with phylogeography in the 2000s, and revived again by geneticists and Kristiansen’s workgroup based on the famous 2015 papers, whereby – due to the “Yamnaya ancestral component” – the Yamna culture would have been composed of communities of R1a-M417 and R1b-M269 lineages which remained against all odds ‘related but separated’ for more than two thousand years, sharing a common unitary language (why? and how?), and which expanded from Yamna (mainly R1b-L23) into Corded Ware (mainly R1a-M417) and then into Bell Beaker (mainly R1b-L51), in imaginary migration waves whose traces Archaeology has not found, or Anthropology described, before.

While phylogeography (especially the distribution of ancient samples of certain R1b and R1a subclades) was the main genetic aspect I used in combination with Archaeology and Anthropology to challenge the reliability of the “Yamnaya ancestral component” in assessing migrations – and thus Kristiansen’s now-popular-again modified Kurgan model – , my main aim was to prove a recent expansion of Late Proto-Indo-European from the steppe, and a still more recent expansion of a common group of speakers of North-West Indo-European, the language ancestral to Italo-Celtic, Germanic, and probably Balto-Slavic (or ‘Temematic’, the NWIE substrate of Balto-Slavic, according to some linguists).

My arguments serve for this purpose, and modern distributions of haplogroups or admixture are fully irrelevant: I am ready to change my view at any time, regarding the role of any haplogroup, or ancestral component, archaeological data, or anthropological migration model, to the extent that it supports the soundest linguistic model.

proto-indo-european-stages
Stages of Proto-Indo-European evolution. IU: Indo-Uralic; PU: Proto-Uralic; PAn: Pre-Anatolian; PToch: Pre-Tocharian; Fin-Ugr: Finno-Ugric. The period between Balkan IE and Proto-Greek could be divided in two periods: an older one, called Proto-Greek (close to the time when NWIE was spoken), probably including Macedonian, and spoken somewhere in the Balkans; and a more recent one, called Mello-Greek, coinciding with the classically reconstructed Proto-Greek, already spoken in the Greek peninsula (West 2007). Similarly, the period between Northern Indo-European and North-West Indo-European could be divided, after the split of Pre-Tocharian, into a North-West Indo-European proper, during the expansion of Yamna to the west, and an Old European period, coinciding with the formation and expansion of the East Bell Beaker group.

Gimbutas’ old theory of sudden and recent expansion served well to support a real community of Proto-Indo-European speakers, as did later the Yamna -> Corded Ware -> Bell Beaker theory that circulated in the 2000s based on modern phylogeography, and as did later partially Anthony’s updated steppe theory (2007). On the other hand, Kristiansen’s long-lasting connections among north-west Pontic steppe cultures and Globular Amphorae and Trypillian cultures, did not fit well with a close community expanding rapidly – although recent genetic data on Trypillia and Globular Amphorae might be compelling him to improve his migration theory.

So, if data turns out to be not as I expect now, I will reflect that in future versions of the paper. I have no problem saying I am wrong. I have been wrong many times before, and something I am certain is that I am wrong now in many details, and I am going to be in the future.

If, for example, R1b-L23(xZ2105) is demonstrated to come from Hungary and not the steppe (as supported by Balanovsky) or R1a-M417 samples are proved to have expanded with West Yamna settlers (as recently proposed by Anthony, see below the Balto-Slavic question), I would support the same model from a linguistic point of view, but modified to reflect these facts. Or if a direct migration link is found in Archaeology from Yamna to Corded Ware, and from Corded Ware to Bell Beaker (as proposed in the 2015 papers), I will revise that too (again, see the image below). Or, if – as Lazaridis et al. (2017) paper on Minoans and Mycenaeans suggested – the Anatolian hypothesis (that is, one of the multiple ones proposed) turns out to be somehow right, I will support it.

calcolithic-expansion
My map of Late Proto-Indo-European expansion (A Grammar of Modern Indo-European, 2006), following Gimbutas and Mallory.

Haplogroups are the least important aspect of the whole model, they are just another data that has to be taken into account for a throrough explanation of migrations. It has become essential today because of the apparent lack of vision on the part of geneticists, who failed to use them to adjust their findings of admixture with findings of haplogroup expansions, favouring thus a marginal theory of long-lasting steppe expansion instead of the mainstream anthropological models.

Since many of these alternative scenarios seem less and less likely with each new paper, it is probably more efficient to talk about which developments are most likely to challenge my model.

Main points

My main predictions – based mostly on language guesstimates, archaeological cultures, and anthropological models of migration -, even with the scarce genomic data we had, have been proven right until know with new samples from Mathieson et al. (2017) and Olalde et al. (2017), among other papers of this past year. These were my original assumptions:

(1) A Middle Proto-Indo-European expansion defined by the appearance of steppe ancestry + reduction in haplogroup diversity and expansion of (mainly) R1b-M269 and R1b-L23 lineages;

(2) A Late Proto-Indo-European expansion defined by steppe ancestry + reduction in haplogroup diversity and expansion of (mainly) R1b-L23 subclades; and

(3) A North-West Indo-European expansion defined by steppe ancestry + reduction in haplogroup diversity and expansion of (mainly) R1b-L51 subclades.

The expansion of Corded Ware peoples, associated with steppe ancestry + reduction in haplogroup diversity and expansion of (mainly) R1a-Z645 subclades, represents thus a different migration, which is compatible with the different nature of the Corded Ware culture, unrelated to Yamna and without migration waves from one to the other (although there were certainly contacts in neighbouring regions).

As you can see, neither of the 3+1 expansion models imply that no other haplogroup can be found in the culture or regions involved (others have in fact been found, and still the models remain valid): these migrations imply a reduction of haplogroup diversity, and the expansion of certain subclades as is common in population expansions throughout history. While we all accept this general idea, some people have difficulties accepting just those cases not compatible with their dreams of autochthonous continuity.

Nevertheless, there are still voids in genetic investigation.

Controversial aspects

In my humble opinion, these are potential conflict periods and the most likely areas of change for the future of the theory:

1. When and how did R1b-M269 lineages become “chiefs” in the steppe?

Based on scarce data from Khvalynsk, it seems that during the Neolithic there were many haplogroups in the North Pontic and North Caspian steppes. A reduction to R1b-M269 subclades must have happened either just before or (as I support) during (the migrations that caused) the Suvorovo-Novodanilovka expansion among Sredni Stog, probably coinciding also with the expansion (or one of the expansions) of CHG ancestry (and thus the appearance of ‘Steppe component’ in the steppe). My theory was based initially on Anthony’s account and TMRCA of haplogroups of modern populations (both ca. 4200-4000 BC), but recent samples of the Balkans (R1b-M269 and steppe ancestry) seem to trace the population expansion some centuries back.

If my assessment is correct, then modern populations of haplogroup R1b-M269* and R1b-L23* in the Balkans probably reflect that ancient expansion, and samples related to Proto-Anatolian cultures in the Balkans will most likely be of R1b-M269 subclades and R1b-L23*. After admixture in the Balkans, posterior migrations of Anatolian languages into Anatolia might be associated with a different admixture component and haplogroups, we don’t have enough data yet.

If the haplogroup reduction and expansion in Khvalynsk happened later than the Suvorovo-Novodanilovka expansion, then we might find the expansion of Pre- or Proto-Anatolian associated with many different haplogroups, such as R1b (xM269), R1a, I, J, or G2, and more or less associated with steppe ancestry in the Balkans.

Another reason for finding such variety of haplogroups in ancient samples from the Balkans would be that this Khvalynsk group of “chiefs” traversed – and mixed with – the Sredni Stog population. Nevertheless, if we suppose homogeneity in haplogroups in Khvalynsk during the expansion, a high proportion of different haplogroups explained by admixture with the local population of Sredni Stog would challenge the whole “chief domination” explanation by Anthony, and we would have to return to the “different culture” theory by Rassamakin and potentially an older migration from Khvalynsk. In any case, both researchers show clear links of the Suvorovo-Novodanilovka phenomenon to Khvalynsk, and a differentiation with the surrounding Sredni Stog culture.

A less likely model would support the identification of the whole Eneolithic Pontic-Caspian steppe as a loose Indo-Hittite-speaking community, which would be in my opinion too big a territory and too loose a cultural bond to justify such a long-lasting close linguistic connection. This will probably be the refuge of certain people looking desperately for R1a-IE connections. However, the nature of the western steppe will remain distinct from Late Proto-Indo-European, which must have developed in the Yamna culture, so autochthonous continuity is not on the table anymore, in any case…

suvorovo-novodanilovka-region
Coexistence of the Varna-Gumelniţa culture and the Suvorovo phase of the sceptre-bearer communities. 1 — Fălciu; 2 — Fundeni-Lungoţi; 3 — Novoselskaja; 4 — Suvorovo; 5 — Casimcea; 6 — Kjulevča; 7 — Reka Devnja; 8 — Drama; 9 — Gonova mogila; 10 — Reževo; 11 — geographically separate Decea variant of the sceptre bearer group (after Govedarica, Manzura 2011: Abb. 5, adapted).

2. How did R1a-M417 (and especially R1a-Z645) haplogroups came to dominate over the Corded Ware cultures?

If I am right (again, based on TMRCA of modern populations), then it is precisely at the time of the potential expansion of Proto-Corded Ware from the Dnieper-Dniester forest, forest-steppe, and steppe regions, ca 3300-3000. Furholt’s recent radiocarbon analysis and suggestions of a Lesser Poland origin of the third or A-horizon, on which disparate archaeologists such as Anthony or Klejn rely now, seem to suggest also that Corded Ware was a cultural complex rather than a compact culture reflecting a migration of peoples – similar thus to the Bell Beaker complex.

This cultural complex interpretation of Corded Ware contrasts with the quite homogeneous late samples we have, suggesting clear migration waves in northern Europe, at least at some point in time, so Genomics will be a great tool to ascertain when and from where approximately did Corded Ware peoples expand. Right now, it seems that Eneolithic Ukraine populations are the closest to its origin, so the traditional interpretation of its regional origin by Kristiansen or Anthony remains valid.

3. How was Indo-Iranian adopted by Corded Ware invaders?

This is rather an anthropological question. We need reasonable models of founder effect/cultural diffusion necessary for that to happen – similar to the ones necessary to explain the arrival of N1c subclades into north-east Europe, or the arrival of R1b subclades in Basque/Iberian-speaking regions in south-west Europe. My description of potential events in the eastern steppe – based partially on Anthony – is merely a short sketch. Genomic data is unlikely to offer more than it does today (replacement of haplogroups, and gradually of some steppe component, by late Corded Ware groups in the steppe), but let’s see what new samples can contribute.

As for what some Indians – and other people willing to confront them – are looking for, regarding R1a-M417 and/or Indo-European origins in India, I don’t see the point, we already know a) that the origin of the expansion is in the steppe and b) that Hindu nationalist biggots will not accept results from research that oppose their views. I don’t expect huge surprises there, just more fruitless discussions (fomented by those who live from trolling or conspiracies)…

4. Yamna settlers from Hungary

Anthony’s new theory – and the nature of Balto-Slavic – hinges on the presence of R1a-M417 subclades (associated with later Corded Ware samples) in Yamna settlers of Hungary, potentially originally from the North Pontic area, where the oldest sample has been found.

My ‘modified’ version of Anthony’s new model (the only I deem just remotely factible) includes the expansion of a Proto-Corded Ware from Lesser Poland, but (given the overwhelming R1b found in East Bell Beaker), with R1a-M417 being associated with the region. How to explain this language change with objective data? Well, we have Bell Beaker expanding to these areas at a later time, so we would need to find R1b-L23 settlers in Lesser Poland, and then a resurge of R1a-M417 haplogroup. If not, resorting yet again to cultural diffusion Yamna “patrons” to Corded Ware “clients” of Lesser Poland would bring us to square one, now with the ‘steppe ancestry’ controversy included…

Since some Eastern Europeans are (for no obvious reason whatsoever) putting their hopes on that IE-R1a-CWC association, let’s hope some samples of R1a-M417 in Yamna or Hungary give them a break, so that they can begin accepting something closer to mainstream anthropological models. We could then work from there a Yamna-> Bell Beaker / North-West Indo-European association truce, and from there keep accepting that no single haplogroup from Yamna settlers is linked with modern languages, cultures or ethnic groups.

yamna-region
localization of Central-European funerary monuments with elements of the Pit Grave culture (after Bátora 2006);

5. How and when was Balto-Slavic associated with haplogroup R1a?

If we accept the Southern or Graeco-Aryan nature of Balto-Slavic with influence from an absorbed North-West Indo-European dialect, “Temematic” (as Kortlandt does), then Indo-Slavonic adopted in the steppe from Potapovka by Sintashta and Poltavka populations divided ca. 2000 BC into Indo-Iranian (migrating to the east with Andronovo), and Balto-Slavic (migrating westward with the Srubna culture). History from there is not straightforward, and it should follow Srubna, Thraco-Cimmerian, or other late expansions from cultures of the steppe.

On the other hand, if it is a Northern dialect related closely to Germanic and Italo-Celtic (in a North-West Indo-European group), then its origin has to be found in the initial expansion of East Bell Beakers, and its development into either the Únětice culture (of Balkan and thus potentially “Southern IE” influence), or the Mierzanowice-Nitra culture (of Corded Ware and thus potentially Uralic influence), or maybe from both, given the intermediate substrate found in Germanic and Balto-Slavic.

It is my opinion that the association of Balto-Slavic with haplogroup R1a is quite early after the East Bell Beaker expansion, probably initially with the subclade typically associated with West Slavic, R1a-M458. I have not much data to support this (apart from the most common linguistic model), just modern haplogroup distribution maps and common TMRCA, and highly hypothetical archaeological-anthropological models. Genetics will hopefully bring more data.

Let’s see also what information on ancient haplogroups we can obtain from the Tollense valley (already showing a close cluster with modern West Slavic populations) and steppe regions.

6. How did Germanic, Celtic, and Italic expand?

Germanic is probably the most interesting one. Following the expansion of R1b-L51 subclades (especially R1b-U106) and steppe ancestry (a confounding factor, with the previous expansion of R1a-Z284 subclades) in Scandinavia is going to be fascinating. Anthropological models already point to a linguistic and archaeological expansion of Pre-Germanic with Bell Beaker peoples.

The expansion of Celtic seems to be associated with chiefdoms, untraceable today in terms of haplogroups, and it seems thus different from previous expansions. New studies might tell how that happened, if it was actually in successive ways, as proposed, or maybe we don’t have enough data yet to reach conclusions.

We don’t know either how Italic expanded into the Italian Peninsula, or whether Latin expanded with peoples from Italy, if at all, or it was mostly a cultural diffusion event, as it seems.

Regarding Etruscan, while I think it is a controversy initiated based on fantastic accounts, and ignited with few finds of Middle Eastern ancestry (that seem logical from the point of view of regional contacts), it will be important for Italian linguists and archaeologists, also to accept the most likely scenario.

As for Palaeo-Hispanic languages, while steppe ancestry is found quite reduced in R1b-L51 subclades (after so many different expansions and admixture events since the departure from the steppe), their distribution from the Chalcolithic onwards and the resurgence of native haplogroups may serve to ascertain which Pre-Roman tribes were associated with the oldest regions where these subclades dominated. For that aim, a closer look at the developments in Aquitania and other pre-Roman Vasconic- and Iberian-speaking regions may shed some light on how founder effects might develop to leave the native language intact (in a case similar to the adoption of Indo-Iranian by post-Corded Ware Sinthastha and Potapovka in the eastern Pontic-Caspian steppe).

NOTE: Although mostly unrelated, linguistic questions may also be somehow altered with a change of migration models. For example, our current Corded Ware Substrate Hypothesis – strongly contested by Kortlandt and others – implies that Uralic was potentially the language spoken by Eneolithic Ukraine / Proto-Corded Ware peoples, therefore early Uralic languages were spoken by Corded Ware peoples, as a substrate for Germanic and Balto-Slavic, and Balto-Slavic and Indo-Iranian. If an Indo-Hittite branch different from Late PIE is accepted for Eneolithic Ukraine (thus suggesting a millennia-long cultural-historical community in the steppe), then the model still stands (e.g. Ger. and BSl. *-mos/-mus, as stated by Kortlandt, would correspond to the oldest morphological IE layer). As you can read in the different versions of our model, the different possibilities for the common substrate are stated, and the most likely one selected. But the most likely a priori option sometimes turns out to be wrong…

NOTE 2: You can comment whatever you want here, but I opened a specific thread in our forum if you want serious comments on the model to stuck and be further discussed.

Featured images: from the book Interactions, changes and meanings. Essays in honour of Igor Manzura on the occasion of his 60th birthday. Țerna S., Govedarica B. (eds.). 2016. Kishinev: Stratum Plus.

See also: