Vikings, Vikings, Vikings! “eastern” ancestry in the whole Baltic Iron Age

Open access Population genomics of the Viking world, by Margaryan et al. bioRxiv (2019), with a huge new sampling from the Viking Age.

Interesting excerpts (emphasis mine, modified for clarity):

To understand the genetic structure and influence of the Viking expansion, we sequenced the genomes of 442 ancient humans from across Europe and Greenland ranging from the Bronze Age (c. 2400 BC) to the early Modern period (c. 1600 CE), with particular emphasis on the Viking Age. We find that the period preceding the Viking Age was accompanied by foreign gene flow into Scandinavia from the south and east: spreading from Denmark and eastern Sweden to the rest of Scandinavia. Despite the close linguistic similarities of modern Scandinavian languages, we observe genetic structure within Scandinavia, suggesting that regional population differences were already present 1,000 years ago.

Maps illustrating the following texts have been made based on data from this and other papers:

  • Maps showing ancestry include only data from this preprint (which also includes some samples from Sigtuna).
  • Maps showing haplogroup density include Vikings from other publications, such as those from Sigtuna in Krzewinska et al. (2018), and from Iceland in Ebenesersdóttir et al. (2018).
  • Maps showing haplogroups of ancient DNA samples based on their age include data from all published papers, but with slightly modified locations to avoid overcrowding (randomized distance approx. ± 0.1 long. and lat.).

middle-ages-europe-y-dna
Y-DNA haplogroups in Europe during the Viking expansions (full map). See other maps from the Middle Ages.

We find that the transition from the BA to the IA is accompanied by a reduction in Neolithic farmer ancestry, with a corresponding increase in both Steppe-like ancestry and hunter-gatherer ancestry. While most groups show a slight recovery of farmer ancestry during the VA, there is considerable variation in ancestry across Scandinavia. In particular, we observe a wide range of ancestry compositions among individuals from Sweden, with some groups in southern Sweden showing some of the highest farmer ancestry proportions (40% or more in individuals from Malmö, Kärda or Öland).

Ancestry proportions in Norway and Denmark on the other hand appear more uniform. Finally we detect an influx of low levels of “eastern” ancestry starting in the early VA, mostly constrained among groups from eastern and central Sweden as well as some Norwegian groups. Testing of putative source groups for this “eastern” ancestry revealed differing patterns among the Viking Age target groups, with contributions of either East Asian- or Caucasus-related ancestry.

saami-ancestry-vikings
Ancestry proportions of four-way models including additional putative source groups for target groups for which three-way fit was rejected (p ≤ 0.01);

Overall, our findings suggest that the genetic makeup of VA Scandinavia derives from mixtures of three earlier sources: Mesolithic hunter-gatherers, Neolithic farmers, and Bronze Age pastoralists. Intriguingly, our results also indicate ongoing gene flow from the south and east into Iron Age Scandinavia. Thus, these observations are consistent with archaeological claims of wide-ranging demographic turmoil in the aftermath of the Roman Empire with consequences for the Scandinavian populations during the late Iron Age.

Genetic structure within Viking-Age Scandinavia

We find that VA Scandinavians on average cluster into three groups according to their geographic origin, shifted towards their respective present-day counterparts in Denmark, Sweden and Norway. Closer inspection of the distributions for the different groups reveals additional complexity in their genetic structure.

vikings-danish-ancestry
Natural neighbor interpolation of “Danish ancestry” among Vikings.

We find that the ‘Norwegian’ cluster includes Norwegian IA individuals, who are distinct from both Swedish and Danish IA individuals which cluster together with the majority of central and eastern Swedish VA individuals. Many individuals from southwestern Sweden (e.g. Skara) cluster with Danish present-day individuals from the eastern islands (Funen, Zealand), skewing towards the ‘Swedish’ cluster with respect to early and more western Danish VA individuals (Jutland).

Some individuals have strong affinity with Eastern Europeans, particularly those from the island of Gotland in eastern Sweden. The latter likely reflects individuals with Baltic ancestry, as clustering with Baltic BA individuals is evident in the IBS-UMAP analysis and through f4-statistics.

vikings-norwegian-ancestry
Natural neighbor interpolation of “Norwegian ancestry” among Vikings.

For more on this influx of “eastern” ancestry see my previous posts (including Viking samples from Sigtuna) on Genetic and linguistic continuity in the East Baltic, and on the Pre-Proto-Germanic homeland based on hydrotoponymy.

Baltic ancestry in Gotland

Genetic clustering using IBS-UMAP suggested genetic affinities of some Viking Age individuals with Bronze Age individuals from the Baltic. To further test these, we quantified excess allele sharing of Viking Age individuals with Baltic BA compared to early Viking Age individuals from Salme using f4 statistics. We find that many individuals from the island of Gotland share a significant excess of alleles with Baltic BA, consistent with other evidence of this site being a trading post with contacts across the Baltic Sea.

vikings-finnish-ancestry
Natural neighbor interpolation of “Finnish ancestry” among Vikings.

The earliest N1a-VL29 sample available comes from Iron Age Gotland (VK579) ca. AD 200-400 (see Iron Age Y-DNA maps), which also proves its presence in the western Baltic before the Viking expansion. The distribution of N1a-VL29 and R1a-Z280 (compared to R1a in general) among Vikings also supports a likely expansion of both lineages in succeeding waves from the east with Akozino warrior-traders, at the same time as they expanded into the Gulf of Finland.

vikings-y-dna-haplogroup-r1a-z280-over-r1a
Density of haplogroup R1a-Z280 (samples in pink) overlaid over other R1a samples (in green, with R1a-Z284 in cyan) among Vikings.

Vikings in Estonia

(…) only one Viking raiding or diplomatic expedition has left direct archaeological traces, at Salme in Estonia, where 41 Swedish Vikings who died violently were buried in two boats accompanied by high-status weaponry. Importantly, the Salme boat-burial predates the first textually documented raid (in Lindisfarne in 793) by nearly half a century. Comparing the genomes of 34 individuals from the Salme burial using kinship analyses, we find that these elite warriors included four brothers buried side by side and a 3rd degree relative of one of the four brothers. In addition, members of the Salme group had very similar ancestry profiles, in comparison to the profiles of other Viking burials. This suggests that this raid was conducted by genetically homogeneous people of high status, including close kin. Isotope analyses indicate that the crew descended from the Mälaren area in Eastern Sweden thus confirming that the Baltic-Mid-Swedish interaction took place early in the VA.

vikings-swedish-ancestry
Natural neighbor interpolation of “Swedish ancestry” among Vikings.

Viking samples from Estonia show thus ancient Swedes from the Mälaren area, which proves once again that hg. N1a-VL29 (especially subclade N1a-L550) and tiny proportions of so-called “Siberian ancestry” expanded during the Early Iron Age into the whole Baltic Sea area, not only into Estonia, and evidently not spreading with Balto-Finnic languages (since the language influence is in the opposite direction, east-west, Germanic > Finno-Samic, during the Bronze Age).

N1a-VL29 lineages spread again later eastwards with Varangians, from Sweden into north-eastern Europe, most likely including the ancestors of the Rurikid dynasty. Unsurprisingly, the arrival of Vikings with Swedish ancestry into the East Baltic and their dispersal through the forest zone didn’t cause a language shift of Balto-Finnic, Mordvinic, or East Slavic speakers to Old Norse, either…

NOTE. For N1a-Y4339 – N1a-L550 subclade of Swedish origin – as main haplogroup of modern descendants of Rurikid princes, see Volkov & Seslavin (2019) – full text in comments below. Data from ancient samples show varied paternal lineages even among early rulers traditionally linked to Rurik’s line, which explains some of the discrepancies found among modern descendants:

  • A sample from Chernihiv (VK542) potentially belonging to Gleb Svyatoslavich, the 11th century prince of Tmutarakan/Novgorod, belongs to hg. I2a-Y3120 (a subclade of early Slavic I2a-CTS10228) and has 71% “Modern Polish” ancestry (see below).
  • Izyaslav Ingvarevych, the 13th century prince of Dorogobuzh, Principality of Volhynia/Galicia, is probably behind a sample from Lutsk (VK541), and belongs to hg. R1a-L1029 (a subclade of R1a-M458), showing ca. 95% of “Modern Polish” ancestry.
  • Yaroslav Osmomysl, the 12th century Prince of Halych (now in Western Ukraine), was probably of hg. E1b-V13, yet another clearly early Slavic haplogroup.

vikings-y-dna-haplogroup-n1a
Density of haplogroup N1a-VL29, N1a-L550 (samples in pink, most not visible) among Vikings. Samples of hg. R1b in blue, hg. R1a in green, hg. I in orange.

Finnish ancestry

Firstly, modern Finnish individuals are not like ancient Finnish individuals, modern individuals have ancestry of a population not in the reference; most likely Steppe/Russian ancestry, as Chinese are in the reference and do not share this direction. Ancient Swedes and Norwegians are more extreme than modern individuals in PC2 and 4. Ancient UK individuals were more extreme than Modern UK individuals in PC3 and 4. Ancient Danish individuals look rather similar to modern individuals from all over Scandinavia. By using a supervised ancient panel, we have removed recent drift from the signal, which would have affected modern Scandinavians and Finnish populations especially. This is in general a desirable feature but it is important to check that it has not affected inference.

ancient-modern-finns-steppe
PCA of the ancient and modern samples using the ancient palette, showing different PCs. Modern individuals are grey and the K=7 ancient panel surrogate populations are shown in strong colors, whilst the remaining M-K=7 ancient populations are shown in faded colors.

The story for Modern-vs-ancient Finnish ancestry is consistent, with ancient Finns looking much less extreme than the moderns. Conversely, ancient Norwegians look like less-drifted modern Norwegians; the Danish admixture seen through the use of ancient DNA is hard to detect because of the extreme drift within Norway that has occurred since the admixture event. PC4 vs PC5 is the most important plot for the ancient DNA story: Sweden and the UK (along with Poland, Italy and to an extent also Norway) are visibly extremes of a distribution the same “genes-mirror-geography” that was seen in the Ancient-palette analysis. PC1 vs PC2 tells the same story – and stronger, since this is a high variance-explained PC – for the UK, Poland and Italy.

Uniform manifold approximation and projection (UMAP) analysis of the VA and other ancient samples.

Evidence for Pictish Genomes

The four ancient genomes of Orkney individuals with little Scandinavian ancestry may be the first ones of Pictish people published to date. Yet a similar (>80% “UK ancestry) individual was found in Ireland (VK545) and five in Scandinavia, implying that Pictish populations were integrated into Scandinavian culture by the Viking Age.

Our interpretation for the Orkney samples can be summarised as follows. Firstly, they represent “native British” ancestry, rather than an unusual type of Scandinavian ancestry. Secondly, that this “British” ancestry was found in Britain before the Anglo-Saxon migrations. Finally, that in Orkney, these individuals would have descended from Pictish populations.

vikings-british-ancestry
Natural neighbor interpolation of “British ancestry” among Vikings.

(…) ‘UK’ represents a group from which modern British and Irish people all receive an ancestry component. This information together implies that within the sampling frame of our data, they are proxying the ‘Briton’ component in UK ancestry; that is, a pre-Roman genetic component present across the UK. Given they were found in Orkney, this makes it very likely that they were descended from a Pictish population.

Modern genetic variation within the UK sees variation between ‘native Briton’ populations Wales, Scotland, Cornwall and Ireland as large compared to that within the more ‘Anglo-Saxon’ English. This is despite subsequent gene flow into those populations from English-like populations. We have not attempted to disentangle modern genetic drift from historically distinct populations. Roman-era period people in England, Wales, Ireland and Scotland may not have been genetically close to these Orkney individuals, but our results show that they have a shared genetic component as they represent the same direction of variation.

Density of haplogroup R1b-L21 (samples in red), overlaid over all samples of hg. R1b among Vikings (R1b-U106 in green, other R1b-L151 in deep red). To these samples one may add the one from Janakkala in south-western Finland (AD ca. 1300), of hg. R1b-L21, possibly related to these population movements.

For more on Gaelic ancestry and lineages likely representing slaves among early Icelanders, see Ebenesersdóttir et al. (2018).

Y-DNA

As in the case of mitochondrial DNA, the overall distribution profile of the Y chromosomal haplogroups in the Viking Age samples was similar to that of the modern North European populations. The most frequently encountered male lineages were the haplogroups I1, R1b and R1a.

Haplogroup I (I1, I2)

The distribution of I1 in southern Scandinavia, including a sample from Sealand (VK532) ca. AD 100 (see Iron Age Y-DNA maps) proves that it had become integrated into the West Germanic population already before their expansions, something that we already suspected thanks to the sampling of Germanic tribes.

vikings-y-dna-haplogroup-i
Density of haplogroup I (samples in orange) among Vikings. Samples of hg. R1b in blue, hg. R1a in green, N1a in pink.
vikings-y-dna-haplogroup-i1-over-i
Density of haplogroup I1 (samples in red) overlaid over all samples of hg. I among Vikings.

Haplogroup R1b (M269, U106, P312)

Especially interesting is the finding of R1b-L151 widely distributed in the historical Nordic Bronze Age region, which is in line with the estimated TMRCA for R1b-P312 subclades found in Scandinavia, despite the known bottleneck among Germanic peoples under U106. Particularly telling in this regard is the finding of rare haplogroups R1b-DF19, R1b-L238, or R1b-S1194. All of that points to the impact of Bell Beaker-derived peoples during the Dagger period, when Pre-Proto-Germanic expanded into Scandinavia.

Also interesting is the finding of hg. R1b-P297 in Troms, Norway (VK531) ca. 2400 BC. R1b-P297 subclades might have expanded to the north through Finland with post-Swiderian Mesolithic groups (read more about Scandinavian hunter-gatherers), and the ancestry of this sample points to that origin.

However, it is also known that ancestry might change within a few generations of admixture, and that the transformation brought about by Bell Beakers with the Dagger Period probably reached Troms, so this could also be a R1b-M269 subclade. In fact, the few available data from this sample show that it comes from the natural harbour Skarsvågen at the NW end of the island Senja, and that its archaeologist thought it was from the Viking period or slightly earlier, based on the grave form. From Prescott (2017):

In 1995, Prescott and Walderhaug tentatively argued that a dramatic transformation took place in Norway around the Late Neolithic (2350 BCE), and that the swift nature of this transition was tied to the initial Indo-Europeanization of southern and coastal Norway, at least to Trøndelag and perhaps as far north as Troms. (…)

The Bell Beaker/early Late Neolithic, however, represents a source and beginning of these institution and practices, exhibits continuity to the following metal age periods and integrated most of Northern Europe’s Nordic region into a set of interaction fields. This happened around 2400 BCE, at the MNB to LN transition.

NOTE. This particular sample is not included in the maps of Viking haplogroups.

vikings-y-dna-haplogroup-r1b
Density of haplogroup R1b (samples in blue) among Vikings. Samples of hg. I in orange, hg. R1a in green, N1a in pink.
vikings-y-dna-haplogroup-r1b-U106-over-r1b
Density of haplogroup R1b-U106 (samples in green) overlaid over all samples of hg. R1b (other R1b-L23 samples in red) among Vikings.
vikings-y-dna-haplogroup-r1b-P312-over-r1b
Density of R1b-L151 (xR1b-U106) (samples in deep red) overlaid over all samples of hg. R1b (R1b-U106 in green, other R1b-M269 in blue) among Vikings.

Haplogroup R1a (M417, Z284)

The distribution of hg. R1a-M417, in combination with data on West Germanic peoples, shows that it was mostly limited to Scandinavia, similar to the distribution of I1. In fact, taking into account the distribution of R1a-Z284 in particular, it seems even more isolated, which is compatible with the limited impact of Corded Ware in Denmark or the Northern European Plain, and the likely origin of R1a-Z284 in the expansion with Battle Axe from the Gulf of Finland. The distribution of R1a-Z280 (see map above) is particularly telling, with a distribution around the Baltic Sea mostly coincident with that of N1a.

vikings-y-dna-haplogroup-r1a
Density of haplogroup R1a (samples in green) among Vikings. Samples of hg. R1b in blue, of hg. I in orange, N1a in pink.
vikings-y-dna-haplogroup-r1a-z284-over-r1a
Density of haplogroup R1a-Z284 (samples in cyan) overlaid over all samples of hg. R1a (in green, with R1a-Z280 in pink) among Vikings.

Other haplogroups

Among the ancient samples, two individuals were derived haplogroups were identified as E1b1b1-M35.1, which are frequently encountered in modern southern Europe, Middle East and North Africa. Interestingly, the individuals carrying these haplogroups had much less Scandinavian ancestry compared to the most samples inferred from haplotype based analysis. A similar pattern was also observed for less frequent haplogroups in our ancient dataset, such as G (n=3), J (n=3) and T (n=2), indicating a possible non-Scandinavian male genetic component in the Viking Age Northern Europe. Interestingly, individuals carrying these haplogroups were from the later Viking Age (10th century and younger), which might indicate some male gene influx into the Viking population during the Viking period.

vikings-italian-ancestry
Natural neighbor interpolation of “Italian ancestry” among Vikings.

As the paper says, the small sample size of rare haplogroups cannot distinguish if these differences are statistically relevant. Nevertheless, both E1b samples have substantial Modern Polish-like ancestry: one sample from Gotland (VK474), of hg. E1b-L791, has ca. 99% “Polish” ancestry, while the other one from Denmark (VK362), of hg. E1b-V13, has ca. 35% “Polish”, ca. 35% “Italian”, as well as some “Danish” (14%) and minor “British” and “Finnish” ancestry.

Given the E1b-V13 samples of likely Central-East European origin among Lombards, Visigoths, and especially among Early Slavs, and the distribution of “Polish” ancestry among Viking samples, VK362 is probably a close description of the typical ancestry of early Slavs. The peak of Modern Polish-like ancestry around the Upper Pripyat during the (late) Viking Age suggests that Poles (like East Slavs) have probably mixed since the 10th century with more eastern peoples close to north-eastern Europeans, derived from ancient Finno-Ugrians:

vikings-polish-ancestry
Natural neighbor interpolation of “Polish ancestry” among Vikings.

Similarly, the finding of R1a-M458 among Vikings in Funen, Denmark (VK139), in Lutsk, Poland (VK541), and in Kurevanikha, Russia (VK160), apart from the early Slav from Usedom, may attest to the origin of the spread of this haplogroup in the western Baltic after the Bell Beaker expansion, once integrated in both Germanic and Balto-Slavic populations, as well as intermediate Bronze Age peoples that were eventually absorbed by their expansions. This contradicts, again, my simplistic initial assessment of R1a-M458 expansion as linked exclusively (or even mainly) to Balto-Slavs.

antiquity-europe-y-dna
Y-DNA haplogroups in Europe during Antiquity (full map). See other maps of cultures and ancient DNA from Antiquity.

Related

Join the discussion...

Please keep the discussion of this post on topic.
For other topics, use the forum instead.
newest oldest most voted
Notify of
trackback

[…] became incorporated into North Eurasian populations of different languages, reaching Uralic- and Indo-European-speaking territories of north-eastern Europe during the late Iron Age, at a time when their ancestral origin […]

trackback

[…] Vikings, Vikings, Vikings! “eastern” ancestry in the whole Baltic Iron Age […]

trackback

[…] Vikings, Vikings, Vikings! “eastern” ancestry in the whole Baltic Iron Age […]

trackback

[…] Vikings, Vikings, Vikings! “eastern” ancestry in the whole Baltic Iron Age […]

Gioiello Tognoni
Gioiello Tognoni

It is good that Carlos Quiles has begun to admit that he was wrong in the past [“This contradicts, again, my simplistic initial assessment of R1a-M458 expansion as linked exclusively (or even mainly) to Balto-Slavs”, p 19], but here he insists in his prejudices. The R-supposed P297 found in 2400 BC Norway is first of all uncertain, given its low coverage [“so this could also be a R1b-M269 subclade” p. 13], and could also be a more recent subclade, but Quiles says that it demonstrates an origin from the Swiderian culture [ “Also interesting is the finding of hg. R1b-P297… Read more »

Tomenable
Tomenable

Anyway one of lessons from this Viking study is that multiculturalism in Scandinavia is nothing new! 😉

Finn Folcwalding
Finn Folcwalding

Not weird because when the Vikings became ‘global’- read European- players the influences went also vice versa. In that range I’m curios about this: Among the ancient samples, two individuals were derived haplogroups were identified as E1b1b1-M35.1, which are frequently encountered in modern southern Europe, Middle East and North Africa. Interestingly, the individuals carrying these haplogroups had much less Scandinavian ancestry compared to the most samples inferred from haplotype based analysis. A similar pattern was also observed for less frequent haplogroups in our ancient dataset, such as G (n=3), J (n=3) and T (n=2), indicating a possible non-Scandinavian male genetic… Read more »

Tomenable
Tomenable

Your map has mistakes, there are also samples with Polish-like DNA from Denmark:

From Supplementary Table S6, samples sorted from most to least of Polish-like DNA:

https://www.biorxiv.org/content/biorxiv/early/2019/07/17/703405/DC3/embed/media-3.xlsx?download=true
comment image

On the other hand the map you posted indicates no Polish-like samples in Denmark:
comment image

Carlos Quiles

Maps show interpolations, and that’s how they work. If there is one or two samples with 90% of something from one region, and a hundred samples from another region where only one or two samples have 100%, you won’t find these “outliers” easily. However, you may notice them because of the weighted areas toward Denmark and Gotland… Maps won’t show whatever it is you are looking for, they show what they show. If it works fine with “British”, “Danish”, “Swedish”, “Norwegian”, “Finnish”, and “Italian” ancestry, and it shows very interesting associations between these modern ancestries and the different Viking expansions,… Read more »

Tomenable
Tomenable

I’m Polish myself and I know the regional genetic profile of every region of Poland (based on my research of Polish samples with confirmed family genealogy from GEDmatch). If the “modern Polish” reference population from this study was based on samples from Western Poles (such as for example Poznań area), then you could probably claim that it has a substantial Germanic component. But as far as I know is based on either Masovians like other studies (North-Eastern Poland) or on Central Poles (Lodz region). And these groups tend to have results closer to Eastern Slavs than to Czechs or East… Read more »

Tomenable
Tomenable

Before speculating that R1a-M458 might be Germanic based on samples from this study, haven’t you noticed (in Supplementary Table S6) that all these R1a-M458 Vikings are autosomally Slavic (“Polish-like” means Slavic), even the one from Funen (where we have also other, non-genetic, evidence of Slavic presence)? Historical records (such as Snorri Sturulson and Saxo Grammaticus), toponymy, archaeology and earlier genetic studies all confirm Slavic/Polish influences in Denmark. For example: 1. This study detected Polish admixture signal in Danish islands dated to before the Middle Ages (and “before the Middle Ages” = before 1050 AD according to Danish chronology): http://eurogenes.blogspot.com/2016/08/on-remarkable-genetic-homogeneity-of.html?m=1 2.… Read more »

Carlos Quiles

I think we all speculate and are bound to be wrong depending on how much data we handle and how we deal with our biases. I can’t know if someone cheated on someone, but if he is a Rurikid and continues the tradition of Norse names, I would say you have to assume a Viking paternal line, even if it’s not from Rurik himself. Anyway, beside those of N1c-L550 subclade, I think you might be right about some other Rurikids being of Slavic lineage in origin. However, about R1a-M458 samples from Eastern Europe being autosomally “Polish”-like, probably like locals, doesn’t… Read more »

Tomenable
Tomenable

As for the tradition of Norse names – Germanic names became very common in Spain after the Visigothic conquest, yet according to Olalde 2019 “The genomic history of the Iberian Peninsula over the past 8000 years”, Visigothic admixture is not that significant in modern Spanish people, and even most of the ancient Visigothic samples published in that study were already mostly non-Germanic autosomally. They were either mixed with the locals, or non-local but non-Germanic either – like I12031. Visigoth I12031 looks like someone who was of Native Balkan-Italian ancestry and was absorbed by the migrating Goths on their way from… Read more »

Carlos Quiles

On Rurikids, which apparently remains a controversial topic for Slavic nation-building myths in the 21st century, we already had reliable information about N1a being one of the main paternal lines among descendants (if not the main one), so the arrival of N1a-L550 subclades with Swedes should not be a surprise. From the Conference Centenary of Human Population Genetics, Moscow, 29-31 May 2019: Genetic study of the Rurik Dynasty Volkov V.G., Seslavin A.N. Tomsk State Pedagogical University, Tomsk, Russia The Rurikids are one of the oldest dynasties of Europe and presently living descendants of Rurik can be found in various locations… Read more »

Tomenable
Tomenable

Right now this is what the evidence suggests:

https://anthrogenica.com/showthread.php?17812-Population-genomics-of-the-Viking-world-(bioxiv-2019-Copenhagen)&p=584819&viewfull=1#post584819

Yaroslav the Wise and Ingegerd had three sons, and their descendants belong to:

Izyaslav – his modern descendants are I2a
Sviatoslav – no descendants survived to present-day (?), but his son Gleb was I2a per this study
Vsevolod – his modern descendants are N1c

^^^
And Vsevolod’s N1c is the suspected product of Ingegerd’s romance with Olaf II Haraldsson.

Here is the theory about Vsevolod being Olaf’s son: https://space-time.ru/space-time/article/view/2226-7271provr_st3-21.2015.71

IMO testing Olaf II Haraldsson’s descendants (or his own bones, if they exist) should follow.

=====

But I understand that “Christian supremacists” will deny that Saint Anna could ever cheat: 😉

https://en.wikipedia.org/wiki/Ingegerd_Olofsdotter_of_Sweden#Sainthood

Joey B&T
Joey B&T

I find the I2a man being Gleb extremely unlikely, as the sample lacked any kind of autosomal Scandinavian input, despite his mother being from Scandinavia. I still think the one lineage thats not found in the local populace but is found almost exlusively in the attested area where the Rus are said to originate from the more likely candidate than the one that could easily be just another local.

But I get it, Slavic posters have been opposing the theory for as long as I remember, no doubt for nationalistic reasons.

Konstantin Demidov
Konstantin Demidov

Not just Rurik. But the rulers of the Grand Duchy of Lithuania – Hediminovichi. Also haplogroup N. And a number of noble names in North-Eastern Europe. In General, the history of North-Eastern Europe of the early middle ages takes place against the background of the struggle of representatives of haplogroup N – no matter how strange it looks. “Nganasans criminal showdown” each other 🙂

Chetan
Chetan

“In fact, taking into account the distribution of R1a-Z284 in particular, it seems even more isolated, which is compatible with the limited impact of Corded Ware in Denmark or the Northern European Plain, and the likely origin of R1a-Z284 in the expansion with Battle Axe from the Gulf of Finland.”

I think one Sarmatian (LS-13 from the southern Urals) from the recent western steppe paper is also shown to be under Z284.

Carlos Quiles

Yes, thank you, I saw that. It’s LS-13, from Novo-Muraptalovo-7, in the northern boundary of the steppe west of the Urals, ca. 484-233 BC. I guess it should be attributed to Eastern Corded Ware groups, even if it hasn’t showed up in the steppe in the previous periods. The problem is its supposed subclade, R1a-S5301, whose modern distribution doesn’t seem to be compatible with an eastern origin, but rather a viking one… It also bothers me that its nomenclature is “LS-” instead of “MJ-” as the rest of samples from the paper, so I would bet maybe for a wrong… Read more »

Konstantin Demidov
Konstantin Demidov

This is not surprising. Most likely the movement of Scandinavian to the East began in the Vandel Period. After all, the main way was not “the way from the Varangians to the Greeks”, but “the Volga-Caspian way”. The campaigns of the Rus in the Caspian sea were described from Ibn Faldan. Although the first campaign Russ dates back to the middle of the 9th century. One of the reasons for this campaign is the ruin of trade factors that ALREADY existed in the Caspian sea.

Chetan
Chetan

“Or did it expand initially with this “eastern” Saami/Sarmatian ancestry of Akozino warrior-traders (like those of hg. N1a-VL29 and R1a-Z280) into Finland, and then into Sweden, and then again westward and eastward with vikings?”

Doesn’t Z284 already occur in Baltic and Nordic CW samples? Although I’m not sure of the subclades. So it’s probably wrongly dated. (seeing how this subclade’s formation date is ~1700 BC). Or like you say, the subclade could have expanded east and west during the Iron Age with the traders.

Carlos Quiles

I have just updated the ancestry maps, because I noticed I failed to include some samples when mixing the sheet of lat/long with the sheet of ancestry. Changes are probably minimal, though.

This post may be particularly slow now, because of the added size of all maps.

I think haplogroup maps are kind of lame, but better-looking alternatives like the map below for R1b-L21 (best suited for “usual” mainland density maps) are actually more artificial in this case.

For seafaring and raiding Vikings, I think those posted above convey a more precise picture for smaller regions and islands.
comment image

Vinitharya
Vinitharya

Yeah they found two L1029’s! It didn’t take a genius, though, to determine at least my subclade YP445 was in the Viking voyages as the first sample I noted was from a John Shanner of Yorkshire, England, a noted area of Danish Viking settlement. Another subclade of L1029 (YP4647, I believe?) also made its way to England with the Batt/Batte family, one descendent of the latter spelling, a Californian, is in my Facebook M458 group.

Carlos Quiles

Yeah, it probably didn’t take a genius, but it certainly took someone with interest in the history of specific subclades…

Anyway, the haplogroup is quite old, so I would stay (as Konstantin Demidov seems to suggest) with a R1a-M458 expanding from the West Baltic, i.e. with Northern Europeans.

Who knows, maybe the Tollense Valley sampling might show an explosion of this haplogroup in the ancient DNA record…

Tomenable
Tomenable

As I wrote in my other response, R1a-M458 is quite old, but its most widespread branches/subclades (and this includes branches in East Germany – because they are generally the same as Polish and Czecho-Slovak branches) usually have young TMRCA. This indicates that during much of its early history M458 was most probably confined geographically to a small area, and probably it was a small population. An alternative explanation is that it could be numerous, but later most of M458 got extinct (population bottleneck). Currently the more likely explanation is the 1st one, because lack of M458 in ancient samples indicates… Read more »

Konstantin Demidov
Konstantin Demidov

And what ancient m458 still known?

Konstantin Demidov
Konstantin Demidov

Very interestingly. The picture of course strange appears. At us in Russia several hundreds of years go fight of “normanist” against “Slavophiles”. The first, based on chronicles, and then and archeology – claim that Russia was founded by Vikings. The second stand on the idea of the autochthonic Slavic population. And the absence argument in a modern gene pool of Russians, Ukrainians, Belarusians of massive crossings with a modern gene pool of Scandinavians was the last argument of Slavophiles-. But it is similar that in the early dark Middle Ages distribution гаплогрупп and the nature of migrations could be other,… Read more »

Stefan
Stefan

I am swedish. Just tested r-z92. As far as i know my fatherline goes back to the 1400. All swedes. I would like to know fron where i got this haplogroup.

trackback

[…] Vikings, Vikings, Vikings! “eastern” ancestry in the whole Baltic Iron Age […]

Tom Simmons
Tom Simmons

Carlos,
I am Z2/Y-18324 (S-22165) with a slow-mutating marker at DYS388 (my value is “10”). I have a number of test kit “matches”, including in the U.K. as well as in Norway and Sweden. Because of the slow-mutating marker, Dr. Iain Mcdonald has estimated that I share a common male ancestor with the Scandinavian kits 1,000+ years ago. If my information can be of value to your very interesting project, please do get in touch.

trackback

[…] Vikings, Vikings, Vikings! “eastern” ancestry in the whole Baltic Iron Age […]