The complexities of 3rd millennium Steppe-related migrations


Open access paper Mobility and Social Change: Understanding the European Neolithic Period after the Archaeogenetic Revolution, by Martin Furholt, J. Archaeol. Res. (2021).

Content under CC-BY license. Interesting excerpts (emphasis mine, stylistic changes for clarity):

This detailed picture of Caucasian population history shows that the initial assertion in the 2015 papers, namely of a one-way migration from east to west, was a simplification supported by a variant of admixture analyses that featured Yamnaya as one unified genetic element (e.g., Haak et al. 2015, fig. 3), which led to calculations of Corded Ware individuals showing 75% Yamnaya ancestry. This

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Earliest R1a-Z93…from Late Trypillia in the Podolian-Volhynian Upland!


Recently, the preprint by Sirak et al. biorXiv (2019), Human auditory ossicles as an alternative optimal source of ancient DNA, was published in Genome Res. (2020), and the corresponding samples were finally uploaded to ENA.

I have been trying to get my hands on sample GLAV_14, a male from the Late Eneolithic site Glăvăneştii Vechi, classified as Romania Bronze Age (ca. 3500-3000 BC), mtDNA T1a1, referenced as investigated first in the study:

Haas N, Maximilian K. 1958. Anthropological study of the human bones from graves with ochre from Glăvăneștii Vechi, Corlăteni and Stoicani Cetățuie. Soviet Anthropology 4,

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On the Ukraine Eneolithic outlier I6561 from Alexandria


Over the past week or so, since the publication of new Corded Ware samples in Narasimhan, Patterson et al. (2019) and after finding out that the R1a-M417 star-like phylogeny may have started ca. 3000 BC, I have been ruminating the relevance of contradictory data about the Ukraine_Eneolithic_o sample from Alexandria, its potential wrong radiocarbon date, and its implications for the Indo-European question.

How many other similar ‘controversial’ samples are there which we haven’t even considered? And what mechanisms are in place to control that the case of Hajji_Firuz_CA I2327 is not repeated?

Ukraine Eneolithic outlier I6561

It was not … Read the rest “On the Ukraine Eneolithic outlier I6561 from Alexandria”

Yamnaya ancestry: mapping the Proto-Indo-European expansions


The latest papers from Ning et al. Cell (2019) and Anthony JIES (2019) have offered some interesting new data, supporting once more what could be inferred since 2015, and what was evident in population genomics since 2017: that Proto-Indo-Europeans expanded under R1b bottlenecks, and that the so-called “Steppe ancestry” referred to two different components, one – Yamnaya or Steppe_EMBA ancestry – expanding with Proto-Indo-Europeans, and the other one – Corded Ware or Steppe_MLBA ancestry – expanding with Uralic speakers.

The following maps are based on formal stats published in the papers and supplementary materials from 2015 until today, mainly on … Read the rest “Yamnaya ancestry: mapping the Proto-Indo-European expansions”

“Steppe ancestry” step by step: Khvalynsk, Sredni Stog, Repin, Yamna, Corded Ware


Wang et al. (2018) is obviously a game changer in many aspects. I have already written about the upcoming Yamna Hungary samples, about the new Steppe_Eneolithic and Caucasus Eneolithic keystones, and about the upcoming Greece Neolithic samples with steppe ancestry.

An interesting aspect of the paper, hidden among so many relevant details, is a clearer picture of how the so-called Yamnaya or steppe ancestry evolved from Samara hunter-gatherers to Yamna nomadic pastoralists, and how this ancestry appeared among Proto-Corded Ware populations.

Please note: arrows of “ancestry movement” in the following PCAs do not necessarily represent physical Read the rest ““Steppe ancestry” step by step: Khvalynsk, Sredni Stog, Repin, Yamna, Corded Ware”

Scythians in Ukraine, Natufian and sub-Saharan ancestry in North Africa (ISBA 8, 21st Sep)


Interesting information from ISBA 8 sesions today, as seen on Twitter (see programme in PDF, and sessions from the 19th and the 20th september).

Official abstracts are listed first (emphasis mine), then reports and images and/or link to tweets. Here is the list for quick access:

Scythian population genetics and settlement patterns

Genetic continuity in the western Eurasian Steppe broken not due to Scythian dominance, Read the rest “Scythians in Ukraine, Natufian and sub-Saharan ancestry in North Africa (ISBA 8, 21st Sep)”

On the origin and spread of haplogroup R1a-Z645 from eastern Europe


In my recent post about the origin and expansion of haplogroup R1b-L51, Chetan made an interesting comment on the origin and expansion of R1a-Z645. Since this haplogroup is also relevant for European history and dialectal North-West Indo-European and Indo-Iranian expansion, I feel compelled to do a similar post, although the picture right now is more blurry than that of R1b-L51.

I find it interesting that many geneticists would question the simplistic approach to the Out of Africa model as it is often enunciated, but they would at the same time consider the current simplistic model of Yamna expansionRead the rest “On the origin and spread of haplogroup R1a-Z645 from eastern Europe”

The Yampil Barrow complex and the Yamna connection with forest-steppe cultures


Researchers involved in the investigation of the Yampil Barrow Complex are taking the opportunity of their latest genetic paper to publish and upload more papers in

NOTE. These are from the free volume 22 of Baltic-Pontic Studies, Podolia “Barrow Culture” Communities: 4th/3rd-2nd Mill. BC. The Yampil Barrow Complex: Interdisciplinary Studies, whose website gives a warning depending on your browser (because of the lack of secure connection). Here is a link to the whole PDF.

Here are some of them, with interesting excerpts (emphasis mine):

1. Kurgan rites in the Eneolithic and Early Bronze age Podolia in Read the rest “The Yampil Barrow complex and the Yamna connection with forest-steppe cultures”

Mitogenomes show likely origin of elevated steppe ancestry in neighbouring Corded Ware groups


Open Access Mitochondrial genomes reveal an east to west cline of steppe ancestry in Corded Ware populations, by Juras et al., Scientific Reports (2018) 8:11603.

Interesting excerpts (emphasis mine, references have been deleted for clarity):

Ancient DNA was extracted from the Corded Ware culture individuals excavated in southeastern Poland (N = 12) and Moravia (N = 3). Late Eneolithic (N = 5) and Bronze Age human remains (N = 25) originated from western Ukraine and came from the Yampil barrow cemetery complex located in the north–western region of the Black Sea. Bronze Age individuals were associated with different archaeological

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