Arrival of steppe ancestry with R1b-P312 in the Mediterranean: Balearic Islands, Sicily, and Iron Age Sardinia

steppe-balearic-sicily-sardinia

New preprint The Arrival of Steppe and Iranian Related Ancestry in the Islands of the Western Mediterranean by Fernandes, Mittnik, Olalde et al. bioRxiv (2019)

Interesting excerpts (emphasis in bold; modified for clarity):

Balearic Islands: The expansion of Iberian speakers

Mallorca_EBA dates to the earliest period of permanent occupation of the islands at around 2400 BCE. We parsimoniously modeled Mallorca_EBA as deriving 36.9 ± 4.2% of her ancestry from a source related to Yamnaya_Samara; (…). We next used qpAdm to identify “proximal” sources for Mallorca_EBA’s ancestry that are more closely related to this individual in space and time, and found that she can be modeled as a clade with the (small) subset of Iberian Bell Beaker culture associated individuals who carried Steppe-derived ancestry (p=0.442).

Suppl. Materials: The model used was with Bell_Beaker_Iberia_highsteppe, a group of outliers from Iberia buried in a Bell Beaker mortuary context who unlike most individuals from this context in that region had high proportions of Steppe ancestry (p=0.442).

Our estimates of Steppe ancestry in the two later Balearic Islands individuals are lower than the earlier one: 26.3 ± 5.1% for Formentera_MBA and 23.1 ± 3.6% for Menorca_LBA, but the Middle to Late Bronze Age Balearic individuals are not a clade relative to non-Balearic groups. Specifically, we find that f4(Mbuti.DG, X; Formentera_MBA, Menorca_LBA) is positive when X=Iberia_Chalcolithic (Z=2.6) or X=Sardinia_Nuragic_BA (Z=2.7). While it is tempting to interpret the latter statistic as suggesting a genetic link between peoples of the Talaiotic culture of the Balearic islands and the Nuragic culture of Sardinia, the attraction to Iberia_Chalcolithic is just as strong, and the mitochondrial haplogroup U5b1+16189+@16192 in Menorca_LBA is not observed in Sardinia_Nuragic_BA but is observed in multiple Iberia_Chalcolithic individuals. A possible explanation is that both the ancestors of Nuragic Sardinians and the ancestors of Talaiotic people from the Balearic Islands received gene flow from an unsampled Iberian Chalcolithic-related group (perhaps a mainland group affiliated to both) that did not contribute to Formentera_MBA.

This sample, like another one in El Argar, is of hg. R1b-P312. So there you are, the data that connects the Proto-Iberian expansion (replacing IE-speaking Bell Beakers) to the Iberian Chalcolithic population, signaled by the increase in Iberian Chalcolithic ancestry after the arrival of Bell Beakers, most likely connected originally to the Argaric and post-Argaric expansions during the MBA.

balearic-sicily-sardinia-pca
PCA with previously published ancient individuals (non-filled symbols), projected onto variation from present-day populations (gray squares).

Steppe in Sardinia IA: Phocaeans from Italy?

Most Sardinians buried in a Nuragic Bronze Age context possessed uniparental haplogroups found in European hunter-gatherers and early farmers, including Y-haplogroup R1b1a[xR1b1a1a] which is different from the characteristic R1b1a1a2a1a2 spread in association with the Bell Beaker complex. An exception is individual I10553 (1226-1056 calBCE) who carried Y-haplogroup J2b2a, previously observed in a Croatian Middle Bronze Age individual bearing Steppe ancestry, suggesting the possibility of genetic input from groups that arrived from the east after the spread of first farmers. This is consistent with the evidence of material culture exchange between Sardinians and mainland Mediterranean groups, although genome-wide analyses find no significant evidence of Steppe ancestry so the quantitative demographic impact was minimal.

Another interesting data, these (Mesolithic) remnant R1b-V88 lineages closely related to the Italian Peninsula, the most likely region of expansion of these lineages into Africa, in turn possibly connected to the expansion of Proto-Afroasiatic.

We detect definitive evidence of Iranian-related ancestry in an Iron Age Sardinian I10366 (391-209 calBCE) with an estimate of 11.9 ± 3.7.% Iran_Ganj_Dareh_Neolithic related ancestry, while rejecting the model with only Anatolian_Neolithic and WHG at p=0.0066 (Supplementary Table 9). The only model that we can fit for this individual using a pair of populations that are closer in time is as a mixture of Iberia_Chalcolithic (11.9 ± 3.2%) and Mycenaean (88.1 ± 3.2%) (p=0.067). This model fits even when including Nuragic Sardinians in the outgroups of the qpAdm analysis, which is consistent with the hypothesis that this individual had little if any ancestry from earlier Sardinians.

yamnaya-samara
Proportions of ancestry using a distal qpAdm framework on an individual basis (a), and based on qpWave clusters

Sicily EBA: The Lusitanian/Ligurian connection?

(…) While a previously reported Bell Beaker culture-associated individual from Sicily had no evidence of Steppe ancestry, (…) we find evidence of Steppe ancestry in the Early Bronze Age by ~2200 BCE. In distal qpAdm, the outlier Sicily_EBA11443 is parsimoniously modeled as harboring 40.2 ± 3.5% Steppe ancestry, and the outlier Sicily_EBA8561 is parsimoniously modeled as harboring 23.3 ± 3.5% Steppe ancestry. (…) The presence of Steppe ancestry in Early Bronze Age Sicily is also evident in Y chromosome analysis, which reveals that 4 of the 5 Early Bronze Age males had Steppe-associated Y-haplogroup R1b1a1a2a1a2. (Online Table 1). Two of these were Y-haplogroup R1b1a1a2a1a2a1 (Z195) which today is largely restricted to Iberia and has been hypothesized to have originated there 2500-2000 BCE. This evidence of west-to-east gene flow from Iberia is also suggested by qpAdm modeling where the only parsimonious proximate source for the Steppe ancestry we found in the main Sicily_EBA cluster is Iberians.

What’s this? An ancestral connection between Sicel Elymian and Galaico-Lusitanian or Ligurian (based on an origin in NE Iberia)? Impossible to say, especially if the languages of these early settlers were replaced later by non-Indo-European speakers from the eastern Mediterranean, and by Indo-European speakers from the mainland closely related to Proto-Italic during the LBA, but see below.

Regarding the comment on R1b-Z195, it is associated with modern Iberians, as DF27 in general, due to founder effects beyond the Pyrenees. It is a very old subclade, split directly from DF27 roughly at the same time as it split from the parent P312, i.e. it can be found anywhere in Europe, and it almost certainly accompanied the expansion of Celts from Central Europe under the subclade R1b-M167/SRY2627.

The connection is thus strong only because of the qpAdm modeling, since R1b-DF27 and subclade R1b-Z195 are certainly lineages expanded quite early, most likely with Yamna settlers in Hungary and East Bell Beakers.

In this case, if stemming from Iberia, it is most likely of subclade R1b-Z220 – or another Z195 (xM167) lineage – originally associated with the Old European substrate found in topo-hydronymy in Iberia, whose most likely remnants attested during the Iron Age were Lusitanians.

r1b-df27-z195
Left: Modern distribution of R1b-Z195 (YFull estimate 2700 BC); Right: Modern distribution of DF27. Both include later founder effects within Iberia, so the increase in the Basque country and the Crown of Aragon and the decrease in Portugal can safely be ignored. Contour maps of the derived allele frequencies of the SNPs analyzed in Solé-Morata et al. (2017).

We detect Iranian-related ancestry in Sicily by the Middle Bronze Age 1800-1500 BCE, consistent with the directional shift of these individuals toward Mycenaeans in PCA. Specifically, two of the Middle Bronze Age individuals can only be fit with models that in addition to Anatolia_Neolithic and WHG, include Iran_Ganj_Dareh_Neolithic. The most parsimonious model for Sicily_MBA3125 has 18.0 ± 3.6% Iranian-related ancestry (p=0.032 for rejecting the alternative model of Steppe rather than Iranian-related ancestry), and the most parsimonious model for Sicily_MBA has 14.9 ± 3.9% Iranian-related ancestry (p=0.037 for rejecting the alternative model).

The modern southern Italian Caucasus-related signal identified in Raveane et al. (2018) is plausibly related to the same Iranian-related spread of ancestry into Sicily that we observe in the Middle Bronze Age (and possibly the Early Bronze Age).

The non-Indo-European Sicanians and Elymians were possibly then connected to eastern Mediterranean groups before the expansion of the Sea Peoples.

For the Late Bronze Age group of individuals, qpAdm documented Steppe-related ancestry, modeling this group as 80.2 ± 1.8% Anatolia_Neolithic, 5.3 ± 1.6% WHG, and 14.5 ± 2.2% Yamnaya_Samara. Our modeling using sources more closely related in space and time also supports Sicily_LBA having Minoan-related ancestry or being derived from local preceding populations or individuals with ancestries similar to those of Sicily_EBA3123 (p=0.527), Sicily_MBA3124 (p=0.352), and Sicily_MBA3125 (p=0.095).

This increase in Steppe-related ancestry in a western site during the LBA most likely represents either an expansion from the Aegean or – maybe more likely, given the archaeological finds – a regional population similar to Sicily EBA re-emerging or rather being displaced from the eastern part of the island because of a westward movement from nearby Calabria.

Whether this population sampled spoke Indo-European or not at this time is questionable, since the Iron Age accounts show non-IE Elymians in this region.

Actually, Elymians seem to have spoken Indo-European, which fits well with the increase in steppe ancestry.

EDIT (21 MAR): Interesting about a proposed incoming Minoan-like ancestry is the potential origin of the Iran Neolithic-related ancestry that is going to appear in Central Italy during the LBA. This could then be potentially associated with Tyrsenians passing through the area, although the traditional description may be more more compatible with an arrival of Sea Peoples from the Adriatic.

Sad to read this:

This manuscript is dedicated to the memory of Sebastiano Tusa of the Soprintendenza del Mare in Palermo, who would have been an author of this study had he not tragically died in the crash of Ethiopia Airlines flight 302 on March 10.

Related

Sintashta diet and economy based on domesticated animal products and wild resources

indo-iranian-sintashta-uralic-migrations

New paper (behind paywall) Bronze Age diet and economy: New stable isotope data from the Central Eurasian steppes (2100-1700 BC), by Hanks et al. J. Arch. Sci (2018) 97:14-25.

Interesting excerpts (emphasis mine):

Previous research at KA-5 was carried out by A. V. Epimakhov in 1994–1995 and 2002–2003 and resulted in the excavation of three Sintashta culture barrows (kurgans) that produced 35 burial pits and a reported 100 skeletons (Epimakhov, 2002, 2005; Epimakhov et al., 2005; Razhev and Epimakhov, 2004). Seven AMS radiocarbon dates on human remains from the cemetery yielded a date range of 2040–1730 cal. BC (2 sigma), which placed the cemetery within the Sintashta phase of the regional Bronze Age (Hanks et al., 2007). Twelve recently obtained AMS radiocarbon dates, taken from short-lived wood and charcoal species recovered from the Kamennyi Ambar settlement, have provided a date range of 2050–1760 cal. BC (2 sigma). Importantly, these dates confirm the close chronological relationship between the settlement and cemetery for the Middle Bronze Age phase and discount the possibility of a freshwater reservoir effect influencing the earlier dating of the human remains from the Kamennyi Ambar 5 cemetery (Epimakhov and Krause, 2013).

Sintashta cemeteries frequently yield fewer than six barrow complexes and the number of skeletons recovered represents a fraction of the total population that would have inhabited the settlements (Judd et al., 2018; Johnson and Hanks, 2012). Scholars have suggested that only members of higher status were afforded interment in these cemeteries and that principles of social organization structured placement of individuals within central or peripheral grave pits (Fig. 2) (Koryakova and Epimakhov, 2007: 75–81). In comparison with other Sintashta cemeteries that have been excavated, KA-5 provides one of the largest skeletal inventories currently available for study.

kamenniy-ambar
Upper – plan of Kamennyi Ambar settlement and cemetery; Lower – plan views of Kurgan 2 and Kurgan 4 from KA-5 Cemetery (kurgan plans redrawn from Epimakhov, 2005: 10, 79).

The KA-5 (MBA), Bestamak (MBA) and Lisakovsk (LBA) datasets exhibited a wide range of δ13C and δ15N values for both humans and herbivores (Figs. 5 and 6 & Table 8). This diversity in isotopic signals may be evident for a variety of reasons. For example, the range of values may be associated with a broad spectrum of C3 and C4 plant diversity in the ancient site biome or herbivore grazing patterns that included more diverse environmental niche areas in the microregion around the sampled sites. Herders also may have chosen to graze animals in niche areas due to recognized territorial boundaries between settlements and concomitant patterns of mobility. Importantly, data from Bolshekaragansky represents humans with lower δ15N values that are more closely associated with δ15N values of the sampled domestic herbivores (Fig. 6). When the archaeological evidence from associated settlement sites is considered, Bolshekaragansky, Bestamak, Lisakovsk and KA-5 have been assumed to represent populations that shared similar forms of pastoral subsistence economies with significant dietary reliance upon domesticated herbivore meat and milk. Human diets have δ13C values closely related to those of local herbivores in terms of the slope of the trendline and range of values (Fig. 6). Comparatively, the cemetery of Bolshekaragansky (associated with the Arkaim settlement) reflects individuals with trend lines closer to those of cattle and caprines and may indicate a stronger reliance on subsistence products from these species with less use of wild riverine and terrestrial resources. The site of Čiča is significantly different with elevated human δ15N isotopic values and depleted δ13C values indicative of a subsistence regime more closely associated with the consumption of freshwater resources, such as fish. The stable isotopic data in this instance is strongly supported by zooarchaeological evidence recovered from the Čiča settlement and also is indicative of significant diachronic changes from the LBA phases through the Iron Age (Fig. 6).

kamenniy-ambar-isotopic-chicha-lisakovsk-bestamark
Regional analysis and comparison of stable isotope results from humans (adults) and animals recovered from MBA and LBA cemeteries in the Southern Urals (Kamennyi Ambar 5 & Bolshekaragansky) northwestern Kazakhstan (Liskovsk & Bestamak) and southwestern Siberia (Čiča).

Conclusion

(…) The isotopic results from KA-5, and recent botanical and archaeological studies from the Kamennyi Ambar settlement, have not produced any evidence for the production or use of domesticated cereals. While this does not definitively answer the question as to whether Sintashta populations engaged in agriculture and/or utilized agricultural products, it does call into serious question the ubiquity of such practices across the region and correlates well with recent archaeological, bioarchaeological, and isotopic studies of human and animal remains from the Southwestern Urals region and Samara Basin (Anthony et al., 2016; Schulting and Richards, 2016). The results substantiate a broader spectrum subsistence diet that in addition to the use of domesticated animal products also incorporated wild flora, wild fauna and fish species. These findings further demonstrate the need to draw on multiple methods and datasets for the reconstruction of late prehistoric subsistence economies in the Eurasian steppes. When possible, this should include datasets from both settlements and associated cemeteries.

Variability in subsistence practices in the central steppes region has been highlighted by other scholars and appears to be strongly correlated with local environmental conditions and adaptations. More comprehensive isotopic studies of human, animal and fish remains are of fundamental importance to achieve more robust and empirically substantiated reconstructions of local biomes and to aid the refinement of regional and micro-regional economic subsistence models. This will allow for a fuller understanding of key diachronic shifts within dietary trends and highlight regional variation of such practices. Ultimately, this will more effectively index the diverse social and environmental variables that contributed to late prehistoric lifeways and the economic strategies employed by these early steppe communities.

Social organization of Sintashta-Petrovka

Interesting to remember now the recent article by Chechushkov et al. (2018) about the social stratificaton in Sintashta-Petrovka, and how it must have caused the long-lasting, peaceful admixture process that led to the known almost full replacement of R1b-L23 (mostly R1b-Z2103) by R1a-Z645 (mostly R1a-Z93) subclades in the North Caspian steppe, coinciding with the formation of the Proto-Indo-Iranian community and language (read my thoughts on this after Damgaard et al. 2018).

Here is another relevant excerpt from Chechushkov et al. (2018), translated from Russian:

settlement-kamenniy-ambar
The map of the settlement of Kamennyi Ambar with excavations, soil cores, and test pits. Legend: a — cuts of the sides of ravines; b — test pits of 2015—2017; c — test pits of 2004; d — soil-science samples with a cultural layer; e — soil-science samples without cultural layer; f — borders of archaeological sites (interpretation of the plan of magnetic anomalies); g — boundaries of excavated structures (1, 2, 4, 5, 7 — Sintashta-Petrovka culture; 3, 6 — Srubnaya-Alakul’ culture).

The analysis suggests that the Sintashta-Petrovka societies had a certain degree of social stratification, expressed both in selective funeral rituals and in the significant difference in lifestyle between the elite and the immediate producers of the product. The data obtained during the field study suggest that the elite lived within the fortifications, while a part of the population was outside their borders, on seasonal sites, and also in stationary non-fortified settlements. Probably, traces of winter settlements can be found near the walls, while the search for summer ones is a task of a separate study. From our point of view, the elite of the early complex societies of the Bronze Age of the Eurasian steppe originated as a response to environmental challenges that created risks for cattle farming. The need to adapt the team to the harsh and changing climatic conditions created a precedent in which the settled collectives of pastoralists – hunter-gatherers could afford the content and magnificent posthumous celebration of people and their families who were not engaged in the production or extraction of an immediate product. In turn, representatives of this social group directed their efforts to the adoption of socially significant decisions, the organization of collective labor in the construction of settlement-shelters and risked their lives, acting as military leaders and fighters.

Thus, in Bronze Age steppe societies, the formation, development and decline of social complexity are directly related to the intensity of pastoralism and the development of new territories, where collectives had to survive in part a new ecological niche. At the same time, some members of the collective took upon themselves the organization of the collective’s life, receiving in return a privileged status. As soon as the conditions of the environment and management changed, the need for such functions was virtually eliminated, as a result of which the privileged members of society dissolved into the general mass, having lost their lifetime status and the right to be allocated posthumously.

Also interesting for the MLBA haplogroup bottleneck in the region is the paper by Judd et al. (2017) about fast life history in Early Indo-Iranian territories.

On the arrival of haplogroup N1c1-L392

Regarding the special position of the Chicha-1 samples in the change of diet and economy during the Iron Age, it is by now well known that haplogroup N must have arrived quite late to North-East Europe, and possibly not linked with the expansion of Siberian ancestry – or linked only with some waves of Siberian ancestry in the region, but not all of them. See Lamnidis et al. (2018) for more on this.

Also, the high prevalence of haplogroup N among Fennic and Siberian (Samoyedic) peoples is not related: while the latter reflects probably the native (Palaeo-Siberian) population that acquired their Uralic branch during the MLBA expansions associated with Corded Ware groups, the former points to the expansion of Fennic peoples into Saamic territory (i.e. after the Fenno-Saamic split) as the most likely period of expansion of N1c1-L392 subclades (see known recent bottlenecks among Finns, and on Proto-Finnic dialectalization).

Probably related to these late incomers are the ancient DNA samples from the Sargat culture during the Iron Age, which show the arrival of N subclades in the region, replacing most – but not all – R1a lineages (see Pilipenko et al. (2017)). Regarding the site of Chicha-1, the following are relevant excerpts about the cultural situation that could have allowed for such stepped, diachronic admixture events in Northern Eurasia, from the paper Stages in the settlement history of Chicha-1: The Results of ceramic analysis, by Molodin et al. (2008):

The stratigraphic data allows us to make the following inference: originally, the settlement was inhabited by people bearing the Late Irmen culture. Later, the people of the Baraba trend of the Suzgun culture arrived at the site (Molodin, Chemyakina, 1984: 40–62). The Baraba-Suzgun pottery demonstrates features similar to what has been reported from the sites of the transitional Bronze to Iron Age culture in the pre-taiga and taiga zones in the Irtysh basin (Potemkina, Korochkova, Stefanov, 1995; Polevodov, 2003). The major morphological types are slightly and well-profiled pots with a short throat. (…)

chicha-irmen-tagar-baraba-forest-siberian
Map showing the location of Chicha-1.

During the following stage of development of the site, the Chicha population increased with people who practiced cultures others than those noted in earlier collections. The ceramic materials from layer 5 provide data on possible relationships. In addition to migrants from northwestern regions practicing the Suzgun culture, there were people bearing the Krasnoozerka culture. Available data also suggests that people from the northern taiga region with the Atlym culture visited the site.

However, people from the west and southwest represent the greatest migration to the region under study. In all likelihood they moved from the northern forest-steppe zone of modern Kazakhstan and practiced the Berlik culture. The spatial distribution analysis of the Chicha-1 site suggests that the Berlik population was rather large. The Berlik people formed a single settlement with the indigenous Late Irmen people and apparently waged certain common economic activities, but preserved their own ethnic and cultural specificity (Molodin, Parzinger, 2006: 49–55). Judging by the data on the chronological sequence of deposited artifacts, migration took place roughly synchronously, hence Chicha-1 became a real cultural and economic center.

(…) In sum, the noted distribution of ceramics over the culture-bearing horizons suggests that beginning with layer 5, traditions of ceramic manufacture described above were practiced, hence the relevant population inhabited the site. Apparently, there were two predominant traditions: the local Late Irmen cultural tradition and the Berlik tradition, which was brought by the immigrants. The Late Irmen people mostly populated the citadel, while the Berlik immigrants inhabited the areas to the east and the north of the citadel.

The stratigraphic data also suggest that the Early Sargat ceramics emerged at the site likely as a part of the Late Irmen tradition (…) Early Sargat ceramics is apparently linked with the Late Irmen tradition. Artifacts associated with the Sargat culture proper have been found in several areas of Chicha-1 (e.g., in excavation area 16). However, the Sargat people appeared at the site after it had been abandoned by its previous inhabitants, and had eventually become completely desolated. This happened no earlier than the 6th cent. BC, possibly in the 5th cent. BC (in fact, the radiocarbon dates for that horizon are close to the turn of the Christian era).

Related

The significance of the Tollense Valley in Bronze Age North-East Germany

bronze-age-tollense-battle

An early Bronze Age causeway in the Tollense Valley, Mecklenburg-Western Pomerania – The starting point of a violent conflict 3300 years ago?, by Jantzen et al. (BERICHT RGK 95, 2014).

Excerpt (emphasis mine):

The causeway in the Tollense Valley, built of timber, stones, turf and sand, and documented over a length of more than 100 m, represents a unique finding from northern Germany. For the first time, part of a Bronze Age network of land routes could be made visible in the southern Baltic area.

Together with the other evidence, the archaeological remains suggest the construction of elaborate trackways and, in some cases, even bridges in the Bronze Age. The Tollense Valley causeway can probably be attributed to the wish or the necessity to be able to cross the Tollense Valley regardless of weather and seasonally differing water level conditions. Its location, situated at a narrow section of the Tollense Valley, offered a prime position for the construction of a permanent crossing of the floodplain on the eastern bank. It is quite possible that a bridge was also part of this.

The complex causeway construction that was likely used and maintained for centuries suggests a significance of the crossing beyond just local. In this context, finds from the valley relating to Bronze Age metal crafts are of interest: along with the scrap metal hoard mentioned above found in the immediate area of the crossing, attention is drawn to a hoard from Golchen comprising an unusual accumulation of tools, as well as to two tin rings found in the same archaeological layer as the Bronze Age skeletal remains. These finds could indicate that metal crafts were of particular significance in the Tollense Valley and its surrounding areas. The middle section of the Tollense Valley that is the focus of attention here could have derived special significance from its role as a crossroads.

The documented pathway, which may have been the starting point of the violent conflict described above, not only contributes to the understanding of the entire findings and the reconstruction of the events in the early 13th century BCE in the Tollense Valley; its context also sheds new light on the cross-regional infrastructure of North-East Germany in the (Early) Bronze Age. Unfortunately, there currently is little further information to integrate it into the broader network of supraregional communication and traffic routes.

The region around the famous barrow of Seddin in Brandenburg is a further example for the significance of river systems for regional power and the exchange of goods. Similarly, the River Tollense could have played a role in the flow of commodities; the causeway at the Kessin 12 site offers a possible connection of the south-north water transportation route via the Tollense River to the Baltic Sea with an east-west land route linking the River Oder estuary region and the Mecklenburg Lake District.

The Lake District was of great importance from the Early Bronze Age; here independent bronze production was established early on. Diversity analyses indicate a shift of regions of innovation during the transition from the 3rd to the 2nd millennium BCE, as the southern Baltic Sea region and the region east of the river Oder clearly also became more important. Early Bronze Age imports from south-east Europe highlight the significance of the region west of the Oder estuary. The Tollense Valley likely played a role in connecting these areas. Therefore, the violent events in the Tollense Valley could also be seen as a result of its strategic significance for the power structure of North-East Germany and the regions on the southern Baltic coast during the Early Bronze Age.

tollensee-valley
Model of the Tollense Valley with the position of pathway (R. Scholz, using a digital model of the valley made by ArcTron [©]).

See also:

The Great Hungarian Plain in a time of change in the Balkans – Neolithic, Chalcolithic, and Bronze Age

hungary-yamna-burials-map

I wrote recently about Anthony’s new model of Corded Ware culture expansion from Yamna settlements of Hungary. I am extremely sceptic about it in terms of current genetic finds, and suspicious of the real reasons behind it – probably misinterpretations of the so-called ‘Yamnaya ancestral component’ in recent genetic papers, rather than archaeological finds.

Nevertheless, it means a definitive rejection by Anthony of:

  • The multiple patron-client relationships he proposed to justify a cultural diffusion of Late Indo-European dialects from Yamna into different Corded Ware cultures in the forest-steppe and Forest Zone (see one of his latest summaries of the model in 2015). Now the language change is explained as a pure migration event, and cultural diffusion is not an option. Ergo, if no migration is found from Hungarian Yamna into Lesser Poland, then Corded Ware cultures were not Indo-European-speaking.
  • Ringe’s glottochronological tree for Proto-Indo-European languages (Ringe, Warnow, and Taylor 2002). An early and sudden split of Late PIE dialects in all directions is substituted by a common, Old European language that expanded from a very small area of settlers, in the Carpathian Basin. This is coincident with the current view on North-West Indo-European, and I think that his final acceptance of a sound linguistic model is essential to solve Indo-European questions.
  • The simplistic assumption of Yamna -> Corded Ware -> Bell Beaker migration found in genetic papers of 2015. The new model implies Yamna->Yamna settlers (Eastern Hungary). Yamna settlers are known to have developed into East Bell Beakers (as described by Gimbutas and accepted by Anthony originally, and now also found in the adoption of Heyd’s theory for his new model); therefore a Yamna settlers (Hungary) -> East Bell Beaker evolution is evident and mainstream, now clear also in genetics. It remains to be seen if the additional Yamna settlers (Hungary) -> Proto-Corded Ware migration proposed by him as a novelty in this new model is also right, i.e. if Yamna settlers from Hungary did in fact migrate into sites of Lesser Poland (to form a Proto-Corded Ware culture). If not, then only Heyd’s model remains.

This new model offers thus a more suitable time frame for usual proto-language guesstimates, that would be compatible with a spread of Late Indo-European with Yamna settlers (of R1b lineages) from the steppe into a small region, where North-West Indo-European would have been spoken, and then a potential cultural diffusion through (or founder effect in) a Proto-Corded Ware culture (of R1a-M417 subclades) of Lesser Poland, which is compatible with the Corded Ware Substrate hypothesis.

Since Anthony has stuck his neck out in favour of this new theory – changing some of his popular theories, and rejecting what many geneticists seem to take as certain – , and because of his previous impressive improvements over Gimbutas’ simple steppe theory (now apparently fashionable again), I think he deserves that his proposal of Yamna/Late Indo-European expansion in the Balkans be further investigated, if only to be improved upon.

I recently found the paper 4000-2000 BC in Hungary: The Age of Transformation, by T. Horváth, in Annales Universitatis Apulensis. Series Historica, 20/II, 51-113. While it deals mainly with the potential survival of the Baden culture into the late third millennium BC, it gives some interesting quite early dates for Yamna (‘Pit’) graves in the Carpathian Basin, and potential cultural (and population) movements within the Balkans.

A note about the Corded Ware culture in the Carpathian Basin:

Many researchers may assume that it is unnecessary for us to deal with the Corded Ware and Globular Amphorae cultures of north Germany, Poland and Denmark, and if so it does not matter what the names of the periods are. It actually matters a lot. It is true that in these areas there was no Baden complex, but the period had many Baden (and other) culture “period phenomena”. These seem to part of a larger formation than cultures – evidenced by traces such as cattle burials, the relationship between copper metallurgies and jade – which link these territories even when the culture complexes were different, because these phenomena appear not just in the Baden, but in the Corded Ware and Globular Amphorae area as well (and these cultural complexes partly overlapped each other both in space and time!). Even the characteristics of sites show many similarities: e.g. in the northern part of corded ware distribution area, mainly burials have been discovered (similarly to the Pit Grave culture in the Great Hungarian Plain) and in the southern part only settlements appear.

At the moment we have no explanation regarding the nature of the relationship between them (it is supposed that as a result of geographical conditions the people of the same culture lived in different ecological conditions and they adapted differently to their environment). In considering the whole of Europe around 3500-3000 BC, easily observable settlement signs disappeared (Milisauskas and Kruk, “Late Neolithic/Late Copper Age,” 307), similarly to Hungary, even though in Hungary this occurred from the end of the Middle Copper Age to the Early Bronze Age, between 4000 and 2000 BC. If we do not take into account that the cattle burials of the Baden culture between 3600 and 2800 BC, and possibly even longer than that, have analogies with the cattle burials of areas in the Early and Middle Neolithic Corded Ware culture (because “logically” analogies would be sought in those areas in the Bronze Age but this period is not analogous with that period in those areas), we would not find any spiritual resemblance in their relationships that lies behind their spatial and temporal analogies; cf. comp. Niels Johannsen and Steffen Laursen, “Routes and Wheeled Transport in Late 4th-Early 3rd Millennium Funerary Customs of the Jutland Peninsula: Regional Evidence and European Context,” PZ 85 (2010): 15-58; Horváth “The Intercultural Connections of the Baden „Culture,” 118. It is painful to think about how many relationships we have not explored or even assessed yet!

hungary-yamna-corded-ware-map
One version from both maps shown in the article, by T. Horváth: “Since the two cultures surely lived together in the Late Copper Age, their collective map represents the Late Copper Age (supplemented with Vučedol sites). Since the direction of diffusion of the Kostolac ceramic style is still unclear, two map versions were made. On one the Kostolac followed the Danube River, on the other they diffused in the opposite direction. In northeast Hungary, Coțofeni III appeared. On this map Kostolac sites are not depicted as dots but, in light of their position and density, proportionately sized arrows are used.”

On Yamna culture and burials in the Carpathian Basin:

Looking at Pit Grave kurgans on the distribution map, it is apparent that burials are the densest where there were no Boleráz or Baden occupations (in this respect this was a kind of “no man’s land”, but from the whole Late Copper Age perspective it was not: the sites of the Baden complex and Pit Grave complemented each other and even partially overlapped). Apart from burials, no Pit Grave settlements or other types of Pit Grave sites are known in Hungary, therefore we do not know whether Pit Grave settlements were situated near the kurgans or whether were somewhere else entirely and we simply have not found them yet.

Since the Pit Grave people had a different lifestyle from the Baden, we can assume that, up to the line of the Tisza River, small animal-keeping mobile groups (Pit Grave) met more populated and settled, agriculturalist, indigenous Boleráz-Baden groups. Animal keepers (Pit Grave) settled in areas where agriculturalists (Boleráz and Baden) did not; in some places, however, they crossed each other’s paths (Fig. 5, 7). Sometimes their sites are very close to each other, sometimes they appear on one site and they can be identified in the stratigraphy of a site. In the latter case the kurgan is always situated on top of a Baden settlement, indicating that Pit Grave not only followed the Baden at these sites but may have represented a somewhat higher social power and belief system than the Baden.

The relationship between pastoral, patrilineal, combatant nomadic tribes and agriculturalist communities is often described as some sort of patron and client relationship. In reality, the signs of such assumption are not visible in the Pit Grave-Baden relationship. There are cases when more aggressive herders conquered more developed agriculturalist communities, but there are also cases when the conqueror’s culture was more developed or stronger than that of the conquered. Always, the conquering nomads are the patrons, the rulers and the empire builders.

In our case, timing is important. How much time had passed on those common sites where a Baden settlement was followed by a Pit Grave kurgan? In these cases, it is certain that the kurgan is younger, but how much younger?

hungary-yamna-settlements
From the article, by T. Horváth. “On the 10 locations analysed, surviving Baden can be assumed after 2800 BC. Unfortunately, it is not possible to predict which sites would survive further scrutiny of radiocarbon dating in this respect; only a few dates are available so far. Therefore, on the map of Baden that still existing after the Late Copper Age, I have also represented all sites (up to the Danube River line) and combined them with Early Bronze Age sites. Since the majority of Makó sites are represented by only one find (scattered finds), and the majority of sites have just one grave, it is impossible to ascertain whether it was part of a cemetery, was within a settlement, or was an individual burial without any further features. Therefore, following Dani 2005, I utilized subdivisions: perhaps in the future this fine subdivision will provide a meaningful explanation (1). Since the radiocarbon dates of Pit Grave kurgans clearly show that the Pit Grave survived at least until 2500 BC, I combined the previous map with that of the Pit Grave. This map would show a realistic picture of cultures after 2800 BC east of the Danube River (2).”

To sum up, the Pit Grave and Baden in the Late Copper Age were certainly contemporary from 3350 BC in the Great Hungarian Plain, and they had common sites, sites which were very close to each other, sites which were far from each other, and also independent sites. The Pit Grave culture surely survived in the transitional period, and into Early Bronze Age I, but perhaps even longer. For the most part, the Baden had ended by 2900 BC in the Great Hungarian Plain. Mapping and some other data (e.g. the discovery that Younger-type, not Mondsee-type, metal objects, which can now be considered to be Baden, even appear east of the Danube River) does not exclude the possibility of searching further for traces of Baden surviving in the Great Hungarian Plain together with or alongside to the Pit Grave. On the common Baden-Pit Grave sites, even without carbon dating, we can assume from already known stratigraphical data that they closely followed each other in time.

For those of you interested in more detailed radiocarbon analysis and assessment of Yamna burials and settlements, from the steppe to the Balkans, to investigate Anthony’s theory further – apart from those authors referenced by him – , I can recommend reading Y. Rassamakin (e.g. Import and Imitation in Archaeology, 2008), S. Ivanova, or Claudia Gerling (e.g. Prehistoric Mobility and Diet in the West Eurasian Steppes 3500 to 300 BC).

Featured image, from the article, by T. Horváth: Distribution map of the Pit Grave.

Related:

The new “Indo-European Corded Ware Theory” of David Anthony

allentoft-yamna-corded-ware

I recently wrote about the Indo-European Corded Ware Theory of Kristian Kristiansen and his workgroup, a sort of “Danish school”, whose aim is to prove a direct, long-lasting interaction between the North Pontic steppe and east European cultures during the Late Neolithic, which supposedly gave rise to a Late Indo-European-speaking Corded Ware culture. That is, a sort of renewed Kurgan model; or, more exactly, Kurgan models, since there is no single one preferred right now.

David Anthony had remained more or less in the background after the controversial assessment of the so-called Yamnaya ancestral component by recent genetic papers, which posited that there was a genetic flow in the Late Neolithic suggesting a migration model that could be hypothetically simplified to Yamna -> Corded Ware -> Bell Beaker.

With his previous publications, especially The Horse, the Wheel, and Language: How Bronze-Age Riders from the Eurasian Steppes Shaped the Modern World (and its revisions), Anthony had set up an impressive revised steppe theory that overcame some of the errors of Gimbutas’ Kurgan model.

Whereas Indo-European-speaking Corded Ware cultures (CWC) were still featured prominently in his model, the different languages supposedly spoken by these groups were explained through multiple cultural diffusion events, and actual migrations from Yamna peoples were only described into Early Bronze Age cultures of the Balkans, and into the Afanasevo culture.

More recently, he has offered (in collaboration with his wife, Dorcas R. Brown) a tentative original connection Yamna -> Corded Ware in the Lesser Poland region, in their paper Molecular archaeology and Indo-European linguistics: Impressions from new data. It seemed to be based merely on recent genetic finds, and on the fact that Corded Ware remains appear to be oldest in that region, according to radiocarbon analysis.

Now he seems to be more and more supportive of this hypothesis in his new essay, Archaeology and Language: Why Archaeologists Care About the Indo-European Problem, in European Archaeology as Anthropology: Essays in Memory of Bernard Wailes ed by P.J. Crabtree and P. Bogucki (2017).

The chapter is interesting to read, as always. Nevertheless, it commits to previous errors – driven by the wrong interpretation of recent genetic papers -, and deepens thus this untenable archaeological-linguistic model of migrations from the steppe, in a weird vicious circle of wrong feedback between archaeologists and geneticists that diregards what archaeologists have been saying in the last decade.

Instead of waiting for the current storm of genetic papers (and their misinformation) to pass, and see what remains, Anthony is now supporting a different model than the one that made him popular, risking the good name he has earned in Archaeology and in popular science texts – in spite of initial setbacks due to the prevailing criticisms of Indo-European migration models.

Some excerpts (emphasis mine):

indo-european-corded-ware-bell-beaker
Central and Eastern Europe ca. 3000–2500 BCE showing the early Yamnaya culture area 3300–2700 BCE and the Yamnaya migration up the Danube Valley with related/offshoot Makó and Vučedol sites; also the distribution of Corded Ware sites in northern Europe; and site areas sampled for aDNA in Haak et al. (2015). The oldest Corded Ware radiocarbon dates are from southern and central Poland. The Yamnaya cemeteries in the Danube Valley are after Heyd (2011), the shaded Globular Amphorae site area is after Harrison and Heyd (2007); the Corded Ware and Globular Amphorae sites in southern Poland are after Machnik (1999); and the blue dots were all Corded Ware sites with radiocarbon dates as of Furholt (2003).

A Yamnaya migration from the steppes up the Danube valley as far as Hungary was already accepted by many archaeologists (Fig. 2.2). Hundreds of Yamnaya-type kurgans and dozens of cemeteries have been recognized by archaeologists in the lower Danube valley, in Bulgaria and Romania; and in the middle Danube valley, in eastern Hungary, with radiocarbon dates that began about 3000–2800 BCE and extended to about 2700–2600 BCE (Ecsedy 1979; Sherratt 1986; Boyadziev 1995; Harrison and Heyd 2007; Heyd 2012; Frînculeasa et al. 2015). The migration stream that created these intrusive cemeteries now can be seen to have continued from eastern Hungary across the Carpathians into southern Poland, where the earliest material traits of the Corded Ware horizon appeared (Furholt 2003). Corded Ware sites appeared in Denmark by 2800–2700 BCE, probably within 100–200 years after the first Yamnaya migrants entered the lower Danube valley. This surprisingly rapid migration introduced genetic traits such as the R1a and R1b Y-chromosome haplogroups and a substantial element of ANE (Ancient North Eurasian) ancestry that remain characteristic of most northern and western Europeans today.

(…)

The oldest radiocarbon dates from Corded Ware sites occur in southern Poland (upper Vistula) and north-central Poland (Kujavia), and this was seen as the region where the early networking of amphorae styles from Globular Amphorae and axe types from Scandinavia began. The genetic evidence shows a somewhat different picture: the Corded Ware people were largely immigrants whose ancestors came from the steppes (probably immediately from eastern Hungary), but they quickly adopted local material traits in amphorae and axe types that obscured their foreign origins. Middle Neolithic northern European populations composed of admixed WHG/EEF survived but were largely excluded from Corded Ware cemeteries, and from marriage into the Corded Ware population. Even centuries after the initial migration the Corded Ware population at Esperstedt, dated 2500–2400 BCE, still exhibited 70–80% Yamnaya genes, although individual variations in the extent of local admixture were apparent. Intermarriage with the surviving local population was more frequent during the ensuing Bell Beaker period. However, the resurgence is more visible in mtDNA than in Y-DNA (Szécsényi-Nagy et al. 2015), suggesting that men of the older EEF heritage were disadvantaged more than women.

(…)

Settlements were more permanent before the Corded Ware migration, and remained so among the Globular Amphorae people, who continued to create more localized site-and-cemetery groups in the same landscape with the more mobile immigrants. Afterward, during the Bell Beaker period, when local genetic ancestry rebounded and the population became more admixed, settlements again were more permanent. The Corded Ware culture introduced both a large, steppe-derived population and an unusually mobile form of pastoral economy that was a regional economic anomaly, but nevertheless survived in varying forms for centuries before the regional economic pattern was re-established. A steppe language certainly accompanied this demographic and economic shift. As we have seen above, there are good independent reasons (loans with Uralic and South Caucasian) to think that PIE was spoken in the steppes. It is likely that the steppe language introduced between 3000–2500 BCE was a late (post-Anatolian) form of PIE and survived and evolved into the later northern IE languages.

So, to sum up the new developments of Anthony’s preferred model:

  1. Abandonment of the multiple cultural diffusion models from Yamna into Corded Ware, i.e. Pre-Germanic (in the Usatovo culture) and Pre-Balto-Slavic (in the Middle Dnieper culture).
  2. The only potential Yamna connection with Corded Ware in Archaeology must come from Yamna migrants in the Carpathian basin. Therefore, R1a must come from Hungarian settlements.
  3. Corded Ware cultures from Northern Europe, from roughly 2800 BC, must come from Yamna settlers of the Carpathian basin.
  4. Esperstedt is a great example of Yamnaya genes, and of the mobility (and lack of intermarriage) of Corded Ware peoples centuries, after their migration from Yamna settlers in Hungary.

My answers (obvious for anyone reading this blog, or my demic diffusion model):

  1. It is a pitty that cultural diffusion models are abandoned. They were the last hope to keep these IE-CWC/Kurgan hypotheses alive.
  2. The Carpathian Basin is obviously the only potential early connection between Corded Ware and Yamna. But no single R1a has been found in western migrants, and admixture (including ancestral components and PCA) from Early Yamna, West Yamna, Balkan EBA, and early Bell Beaker samples from Hungary make it very unlikely that such a connection existed.
  3. Corded Ware peoples formed and began their migration much earlier than Yamna settlers arrived in the Carpathian Basin. Compare e.g. the Late Neolithic sample from Latvia (dated ca. 2885 BC) with steppe ancestry attributed to Corded Ware, or the early appearance of east European cultures like Fatyanovo-Balanovo or Abashevo. Also, known Yamna migration routes don’t include these proposed population expansions.
  4. I have already written about the Esperstedt outlier, and why its definition as an outlier should have been clearly made to avoid this kind of misinterpretations…
yamna-bell-beaker
Yamna – East Bell Beaker migration ca. 3000-2300 BC, according to Heyd (2007)

With each new genetic paper it is less and less likely that many individuals of Y-DNA haplogroup R1a, and especially R1a-Z645 (if any at all), will appear associated with Yamna, either in the Pontic-Caspian steppe or in western settlements (at least clearly belonging to Yamna, Balkan EBA, or Bell Beaker cultures), which will make the life of this new Indo-European Corded Ware Theory model still shorter than could be a priori expected for any archaeological model.

Also, it seems that the Bell Beaker preprint paper by Olalde et al. (2017) will be published in Nature soon with more samples, so a swift rejection of this theory may be near. On the other hand, the first paper on this model by Anthony and Brown (like the first paper of Kristiansen and his workgroup) appeared just before Olalde et al. (2017) and Mathieson et al. (2017), and yet all samples against their pet theories have not deterred any of them to continue supporting them…

I would say it is a shame that some geneticists are misleading good archaeologists into so many different wrong models, but I guess it is only fair to blame authors for what they write, not whom or what they trusted to write…

I think there is much more to be said about the interaction among Neolithic cultures from the steppe (viz. Sredni Stog and Khvalynsk), than about the Yamna migration, and Anthony was in a better position to judge this. Right now, it seems that other researchers like Rassamakin or Ivanova are taking the lead in the research of Neolithic cultures from the steppe, while Heyd or Prescott are taking the lead in the explanation of Yamna -> Bell Beaker migrations and their connection with the expansion of Late Indo-European languages.

#EDIT (December 18 2017): Just to be clear, Anthony’s new Indo-European Corded Ware Theory model in Archaeology would be compatible with the development and expansion of a North-West Indo-European dialect of Late Indo-European in Linguistics (which is my main source of disagreement with other recent models). In fact, Anthony’s new model could explain the different nature of Balto-Slavic, being adopted by peoples of mainly R1a-Z645 subclades of Lesser Poland – from Yamna migrants of R1b-L23 subclades – , and later influencing Pre-Germanic brought by Bell Beakers to Scandinavia, so in that sense it could offer some light to certain controversial linguistic aspects. See Corded Ware Substrate Theory for more on Germanic and Balto-Slavic similarities based on a common, intermediate substrate.

What I am criticising with this post is that the model seems to rely heavily (in fact, almost solely) on what some geneticists (and especially amateurs, fanboys of specific haplogroups and/or admixture components) are selling about the ‘Yamnaya component’ (and thus the assumption of a common migration of peoples of R1a-Z645 and R1b-L23 subclades), something which is – to say the least – highly controversial today. Instead of departing from Archaeology (his field) to try and make sense of what others are saying, he seems to be abandoning his own migration models and adopting one compatible with genetic studies of 2015-2016 made by laymen in Indo-European studies, who based their conclusions on their own new methods, applied to a few scattered samples. These new IECWT proponents are thus in turn giving still more reasons for these geneticists to support wrong assumptions in future studies, by relying on any of these new potential archaeological scenarios. And so on and on it goes…

Related: