The Lusatian culture, the most likely vector of Balto-Slavic expansions

early-bronze-age-languages-europe

New archaeological paper (behind paywall) New evidence on the southeast Baltic Late Bronze Age agrarian intensification and the earliest AMS dates of Lens culinaris and Vicia faba, by Minkevičius et al. Vegetation History and Archaeobotany (2019).

Interesting excerpts (emphasis mine):

Arrival of farming in the south-east Baltic

The current state of research reveals no firm evidence of crop cultivation in the region before the LBA (Piličiauskas et al. 2017b; Grikpėdis and Motuzaitė-Matuzevičiūtė 2018). Current archaeobotanical data firmly suggest the adoption of farming during the EBA to LBA transition. (…) By comparison, in other parts of N Europe subsistence economy of CWC groups was characterized by strong emphasis on animal husbandry, however crop cultivation was also used (Kirleis 2019; Vanhanen et al. 2019). CWC sites from the Netherlands, Denmark, Sweden and Germany reveal evidence of the cultivation of H. vulgare var. nudum, T. dicoccum, Linum usitatissimum (flax) (Oudemans and Kubiak-Martens 2014; Beckerman 2015; Kubiak- Martens et al. 2015).

It is (…) striking that earliest evidence of farming in the SE Baltic only appears in the deposits dating over 4,000 years later.

The environmental conditions of the SE Baltic presented a significant barrier and numerous genetic adaptations were required before farming could successfully spread into the region (Motuzaitė-Matuzevičiūtė 2018). Adaptations through seasonality changes usually play a major role in adapting to new environments (Sherratt 1980). These include establishing genetic controls on seasonality, especially flowering times and length of growing season (Fuller and Lucas 2017). Therefore, it could be argued that farming was only firmly established in the region around the LBA after several crop species, primarily barley, became adapted to the local environment and the risk of crop failure was reduced (Motuzaitė-Matuzevičiūtė 2018). The transition to farming was further aided by the climate warming which started around 750 cal bc (Gaigalas 2004; Sillasoo et al. 2009). In such a case the fragmented evidence from earlier periods is a likely illustration of the early attempts that have failed.

south-east-baltic-agrarian-communities
Map of sites mentioned in the text: 1 Duba and Palesa Lakes, 2 Šventoji, 3 Šarnelė, 4 Iru, 5 Kvietiniai, 6 Kreiči, 7 Turlojiškė, 8 Narkūnai, 9 Luokesa 1, 10 Mūkakalns, 11 Kivutkalns, 12 Asva, 13 Kukuliškiai

Social change

The LBA agrarian intensification of the SE Baltic was most likely not an isolated case but rather a part of broader social, economic and technological developments sweeping across northern Europe.

Evidence from sites across the Baltic Sea shows that the end of the EBA (ca. 1200 bc onward, after Gustafsson 1998) was marked by intensification of agriculture and changes in landscape management. This coincides with the agricultural developments observed on the SE fringes of the Baltic Sea and provides a context for the eventual arrival of farming, followed shortly by the rapid agrarian intensification of the region. Looking just south from the study region, we see that data from northern Poland reveal a sharp increase in both scale and intensity of agricultural activities during the EBA to LBA transition. Pollen records show significant environmental changes starting around 1400/1300 bc (Wacnik 2005, 2009; Wacnik et al. 2012). These were mostly a result of development of a production economy based on plant cultivation and animal raising. Even more significant changes during this period are visible in southern Scandinavia. Pollen records from S Sweden present evidence for an opening up of the forested landscape and the creation of extensive grasslands (Berglund 1991; Gustafsson 1998). Major changes are also apparent in archaeobotanical assemblages.

In general, during the end of the EBA northern Europe underwent a massive transformation of the farming system moving towards a more intensified agriculture aimed at surplus production. However, this should not be regarded as an isolated occurrence, but rather as a radical change of the whole society which took place throughout Europe (Gustafsson 1998). Intensification of contacts across northern Europe have integrated previously isolated regions into a wider network (Kristiansen and Larsson 2005; Wehlin 2013; Earle et al. 2015). It is therefore likely that farming spread into the SE fringes of the Baltic Sea alongside other innovations including malleable technologies and developments of social structure.

bronze-age-late-baltic
Late Bronze Age cultures in the Baltic. See full map.

The presence and scale of intensifying connections is well illustrated by SE Baltic archaeological material.

Firstly, the appearance of stone ship graves has served as a basis for locating the Nordic communication zones. Construction of such graves was limited to the coastal regions of Kurzeme, Saaremaa Island and the Northern Estonian coast near Tallinn and Kaliningrad (Graudonis 1967; Okulicz 1976; Lang 2007) and is generally regarded as a foreign burial custom which was common in Gotland and along the Scandinavian coast. This is also supported by the Staldzene and Tehumardi hoards (Vasks and Vijups 2004; Sperling 2013), which contained artefacts typical of Nordic culture.

Secondly, studies of early metallurgy and its products, both imported and created in the SE Baltic, have concluded that metal consumption in the LBA had more than doubled compared to the EBA (Sidrys and Luchtanas 1999). The SE Baltic region lacks any metal artefact types exclusive to the region and metal objects are dominated by artefact types originating from Nordic and Lusatian cultures (Sidrys and Luchtanas 1999; Lang 2007; Čivilytė 2014). This indicates that even after metal crafting reached the region, the technology remained exclusively of foreign origin. Rarely identifiable negatives of clay casting moulds were also made for artefacts of Nordic influence, such as Mälar type axes or Härnevi type pins (Čivilytė 2014; Sperling 2014).

Lastly, emerging social diversification was accompanied by the establishment of the first identifiable settlement pattern. Settlement locations were strategically chosen alongside economically significant routes, primarily on the coast and near the Daugava River. Hilltop areas were prioritized over the lowlands, and excavations on these sites have often revealed several stages of enclosure construction (Graudonis 1989). This has also been explained as a reflection of intensifying communication networks between Nordic and Lusatian cultures, and the indigenous communities of the SE Baltic.

Proto-Balto-Slavic

One of the aspects of my description of Balto-Slavic I am least convinced about is my acceptance of Kortlandt’s dialectal classification into Proto-East Baltic, Proto-West Baltic, and Proto-Slavic, due to its strong reliance on his own controversial theory of late laryngeal loss.

Kortlandt’s position regarding Balto-Slavic is that it is in fact simply ‘Proto-Baltic’, a language that would stem thus from an Indo-Baltic branch, which would be originally represented by Corded Ware, and which would have split suddenly in its three dialects without any common development between branches, including some intermediate hypothetic “Centum” Temematic substrate that would explain everything his model can’t…

As more genetic and archaeological data on northern Europe appears, his ideas about Balto-Slavic are becoming even less credible, fully at odds with his predicted population and cultural movements, in particular because of the evident shaping of Indo-European-speaking Europe through the expansion of the Bell Beaker culture from the Yamnaya of the Carpathian Basin, and of the shaping of Uralic-speaking Europe through the expansion of the Corded Ware culture.

bronze-age-middle-northern-europe
Middle Bronze Age cultures close to the Baltic ca. 1750-1250 BC. See full map.

The site of Turlojiškė in southern Lithuania (ca. 908-485 BC) – which Mittnik et al. (2018) classified as “Bronze Age, Trzciniec culture?” – can be more reasonably considered a settlement of incoming intensive agrarian communities under the influence of the Lusatian culture, like the Narkūnai hilltop settlement in eastern Lithuania (ca. 800–550 BC), or the enclosed hilltop settlement of Kukuliškiai in western Lithuania (ca. 887-506 BC), just 300 m east of the Baltic Sea, also referred to in the paper.

While the dates of sampled individuals include a huge span (ca. 2100-600 BC), those with confirmed radiocarbon dates are more precisely dated to the LBA-EIA transition. More specifically, the first clearly western influence is seen in the early outlier Turlojiškė1932 (ca. 1230-920 BC), while later samples and samples from Kivutkalns, in Latvia, show major genetic continuity with indigenous populations, compatible with the new chiefdom-based systems of the Baltic and the known lack of massive migrations to the region.

Contacts with western groups of the Nordic Bronze Age and Lusatian cultures intensified – based on existing archaeological and archaeobotanical evidence – in the LBA, especially from ca. 1100/1000 BC on, and Baltic languages seem to have thus little to do with the disappearing Trzciniec culture, and more with the incoming Lusatian influence.

Both facts – more simple dialectalization scheme, and more recent Indo-European expansion to the east – support the spread of Proto-Baltic into the south-east Baltic area precisely around this time, and is also compatible with an internal separation from Proto-Slavic during the expansion of the Lusatian culture.

pca-late-bronze-age-balto-slavic-finnic
Top Left:Likely Baltic, Slavic, and Balto-Finnic-speaking territories (asynchronous), overlaid over Late Bronze Age cultures. Balto-Slavic in green: West(-East?) Baltic (B1), unattested early Baltic (B2), and Slavic (S). Late Balto-Finnic (F) in cyan. In red, Tollense and Turlojiškė sampling. Dashed black line: Balto-Slavic/West Uralic hydrotoponymy border until ca. 1000 AD. Top right: PCA of groups from the Early Bronze Age to the Late Bronze Age. Marked are Iwno/Pre-Trzciniec of Gustorzyn (see below), Late Trzciniec/Iron Age samples from Turlojiškė, and in dashed line approximate extent of Tollense cluster; Y-DNA haplogroups during the Late Bronze Age (Bottom left) and during the Early Iron Age (Bottom right). Notice a majority non-R1a lineages among sampled Early Slavs. See full maps and PCAs.

Even though comparative grammar is traditionally known to be wary of resorting to language contamination or language contact, the truth is that – very much like population genomics – trying to draw a ‘pure’ phylogenetic tree for Balto-Slavic has never worked very well, and the most likely culprit is the Slavic expansion to the south-east into territories which underwent different and complex genetic and linguistic influences for centuries (see here and here).

The close interaction of Nordic BA and Lusatian cultures (and their cultural predominance over) indigenous eastern Baltic peoples from ca. 1100 BC fits (part of) the known intense lexical borrowings of Balto-Finnic from Palaeo-Germanic and from early Proto-Baltic, as well as (part of) the known Germanic–Balto-Slavic contacts, whereas the evident Balto-Finnic-like substrate of Balto-Slavic, and especially of Baltic, must stem from the acculturation of those indigenous East Baltic peoples.

The relative chronology of hydrotoponymy in the East Baltic shows that essentially all ancestral layers to the north of the Daugava must have been Uralic, while roughly south of the Daugava they seem to be mostly Indo-European. The question remains, though, when did this Indo-European layer start?

Despite the many centuries that could separate the attestation of southern place- and river-names from northern ones, Old European is also defined by linguistic traits, which would imply that the same language inferred from Western and Southern European hydrotoponymy is that found in the Baltic, hence all from North-West Indo-European-speaking Bell Beakers and derived Early European Bronze Age groups.

Interestingly, though, it is well-known that some modern Baltic toponyms can’t be easily distinguished from the Old European layers – unlike those of Iberia or the British Isles, which show some attested language change in the proto-historical and historical period – which may imply both (a) continuity of Baltic languages since the EBA, but also that (b) the Baltic naming system is a confounding factor in assessing the ancestral expansion of Old European. The latter is becoming more and more likely with each new linguistic, archaeological, and genetic paper.

up-river
Hydronyms in up-. One among many examples of scarcely attested appellatives that appear inflated in the Baltic due to modern use.

To sum up, a survival of a hypothetical late Trzciniec language in Lithuania or as part of the expanding Lusatian community is not the most economic explanation for what is seen in genetics and archaeology. On the other hand, the cluster formed by the Tollense samples (a site corresponding to the Nordic Bronze Age), the Turlojiškė outlier, and the early Slavs from Bohemia all depict an eastward expansion of Balto-Slavic languages from Central Europe, at the same time as Celtic expanded to the west with the Urnfield culture.

NOTE. Another, more complicated question, though, is if this expanding Proto-Baltic language accompanying agriculture represents the extinct
early Proto-Baltic dialect from which Balto-Finnic borrowed words, hence Proto-Baltic proper expanded later, or if this early Baltic branch could have been part of the Trzciniec expansion. Again, the answer in archaeological and genetic terms seems to be the former. For a more detailed discussion of this and more, see European hydrotoponymy (IV): tug of war between Balto-Slavic and West Uralic.

As I said recently, the slight increase in Corded Ware-like ancestry among Iron Age Estonians, if it were statistically relevant and representative of an incoming population – and not just the product of “usual” admixture with immediate neighbours – need not be from south-eastern Corded Ware groups, because the Akozino-Malär cultural exchange seems to have happened as an interaction in both directions, and not just as an eastward migration imagined by Carpelan and Parpola.

Archaeology and genetics could actually suggest then (at least in part) an admixture with displaced indigenous West Uralic-speaking peoples from the south-west, to the south of the Daugava River, at the same time as the Indo-European – Uralic language frontier must have shifted to its traditional location, precisely during the LBA / EIA transition around 1000 BC.

NOTE. For more on this, see the supplementary materials of Saag et al. (2019).

fortified-settlements-lba-ia
Distribution of fortified settlements (filled circles) and other hilltop sites (empty circles) of the Late Bronze Age and Pre-Roman Iron Ages in the East Baltic region. Tentative area of most intensive contacts between Baltic and Balto-Finnic communities marked with a dashed line. Image modified from (Lang 2016).

The tight relationship of the three communities also accounts for the homogeneous distribution of expanding haplogroup N1c-VL29 (possibly associated with Akozino warrior-traders) in the whole Baltic Sea area, such as those appearing in the Estonian Iron Age samples, which have no clearly defined route(s) of expansion.

It is even possible that they emerged first in the south, linked to marriage alliances of Akozino chieftains with Baltic- and Germanic-speaking chiefdoms around the Baltic Sea (see N1c in Germanic Iron Age), since the expansion of (some) N1c lineages with Gulf of Finland Finnic to the north was more clearly associated with their known bottleneck ca. 2,000 years ago.

Related

European hydrotoponymy (IV): tug of war between Balto-Slavic and West Uralic

germanic-balto-slavic-expansion

In his recent paper on Late Proto-Indo-European migrations, when citing Udolph to support his model, Frederik Kortlandt failed to mention that the Old European hydrotoponymy in northern Central-East Europe evolved into Baltic and Slavic layers, and both take part in some Northern European (i.e. Germanic – Balto-Slavic) commonalities.

Proto-Slavic

From Expansion slavischer Stämme aus namenkundlicher und bodenkundlicher sicht, by Udolph, Onomastica (2016), translated into English (emphasis mine):

NOTE. An archived version is available here. The DOI references for Onomastica do not work.

(…) there is a clear center of Slavic names in the area north of the Carpathians. Among them are root words of the Slavic languages such as reka / rzeka, potok u. a. m.

Even more important than this mapping is the question of how the dispersion of ancient Slavic names happened. What is meant by ancient Slavic names? I elaborated on this in this journal years ago (Udolph, 1997):

(1)Ancient suffixes that are no longer productive today.

This clearly includes Slavic *-(j)ava as in Vir-ava, Vod-ava, Il-ava, Glin-iawa, Breg-ava, Ljut-ava, Mor-ava, Orl-java among others. It has clear links to the ancient common Indo-European language (Lupawa, Morava-March-Moravia, Orava, Widawa). They have a center north of the Carpathians.

ava-slavic

(2) Unproductive appellatives (water words), which have disappeared from the language, are certain witnesses of ancient Slavic settlements. A nice example of this is Ukr. bahno, Pol. bagno ‘swamp, bog, morass’ etc. The word has long been missing in South Slavic, although it appears in South Slavic names, but only in very specific areas (see Udolph, 1979, pp. 324-336).

(3) Names that go back to different sound shifts. [Examples:]

  • (…) the Slavic clan around Old Sorbian brna ‘feces, earth’, Bulgarian OCS brьnije ‘feces, loam’, OCS brъna ‘feces’, Slovenian brn, ‘river mud’, etc. is solved with the inclusion of onomastic materials (Udolph, 1979, p. 499-514). (…) Toponymic mapping shows important details.
  • bryn-slavic
    Karte 4. brъn < *brŭn und bryn- < *brūn- in slavischen Namen
  • (…)We also have an ablauting *krŭn-:*krūn- in front of us. Map 5 shows the distribution of both variants in Slavic names.
  • The next case is quite similar. It concerns Russ. appellative grjaz’ ‘dirt, feces, mud’, (…) for which an Old Slavic form *gręz exists. Slavic also knows the ablauting variant *grǫz.

    These maps (see Map 6, p. 222) show that a homeland of Slavic tribes can only be inferred north of the Carpathians.

    (4) Place-names formed by Slavic suffixes of Pre-Slavic nature, i.e. derived from Old European hydronyms.

    (a) The largest river in Poland, the Wisła, German Vistula, bears a clearly Pre-Slavic name, no matter how one explains it (Babik, 2001, pp. 311-315; Bijak, 2013, p. 34, Udolph, 1990 , Pp. 303-311).

    (b) With the same suffix are formed Sanok, place on the southwest of Przemyśl; Sanoka, a no longer known waters name, 1448 as fluvium Szanoka, near the place Sanoka and with a diminutive suffix -ok- a tributary of the Sanok, which is called Sanoczek (for details see Udolph, 1990, pp. 264-270; Rymut / Majtan, 1998, p. 222). The San also has a single-language name, but that does not change anything about the right etymology. The suffix variant -očь also includes Liwocz and Liwoczka, river names near Cracow; also a mountain range of the Beskydy is mentioned at Długosz as Lywocz.

    According to the opinion of the “Słownik prasłowiański” (Sławski (red.), 1974, p. 92), the suffix -ok- represents a Proto-Slavic archaism. It appears, for example, in sъvědokъ, snubokъ, vidokъ, edok, igrok, inok among others, but its antiquity also shows, among other things, that it started at archaic athematic tribes.

    east-slavic-language-expansion
    Mapping of older and younger East Slavic place-names and translation into settlement evolution.

    Slavonic Urheimat

    If we apply this to the loess distribution in western Ukraine and south-eastern Poland, it is very noticeable that the center of the Old Slavic place names lies in the area where loess dispersal is gradually “frayed out”, i.e. for example, in the area west of Kiev between Krakow in the west and Winnycja and Moldavia in the east. In short, the distribution of good soils coincides with ancient Slavic names. If that is correct, we can expect a homeland in the Pre-Carpathian region, or better, a core landscape of Slavic settlement.

    The existence of Pre-Slavic Indo-European place names and water names whose structure indicates that they originated from an Indo-European basis, but then also developed Slavic peculiarities, can now – as stated above – only be understood to mean that the language group that we call today Slavic emerged in a century-long process from an Indo-European dialectal area.

    Loess areas between Poland and Ukraine. Image from Jary et al. (2018).

    From a genetic point of view, the scarce data published to date show a clear shift of central-east populations from more Corded Ware-like groups in the EBA towards more BBC-derived ancestry in the common era, to the point where ancient DNA samples from East Germany, Poland and Lithuania evolve from clustering between Corded Ware and Sub-Neolithic peoples to clustering close to Bell Beaker-derived groups, such as West Germanic peoples, Tollense samples, etc. (see below)

    Furthermore, sampled Early Slavs show bottlenecks under “Dinaric” I2a-L621 and central-eastern E1b-V13, which – in combination with the known phylogeography of Únětice and Urnfield – is compatible with its late expansion from a central-east European Slavonic homeland, such as the Pomeranian culture, in turn likely derived from Lusatian culture groups.

    This doesn’t preclude a more immediate expansion of Common Slavic in Antiquity closer to the northern Carpathians, which is also supported by the available Early Slavic sampling, apart from samples from the Avar and Hungarian polities.

    pca-balto-slavic-iron-age
    Likely Baltic (yellow-green) and Slavic (orange) groups ca. 500 AD on, with Finnic (cyan) and Mordvinic (blue) groups roughly divided through hydrotoponymy line ca. 1000 AD Top Left: Late Iron Age cultures. Top right: PCA of groups from the Iron Age to the Middle Ages. Y-DNA haplogroups during the Germanic migrations (Bottom left) and during the Middle Ages (Bottom right). Notice a majority non-R1a lineages among sampled Early Slavs. See full maps and PCAs.

    Proto-Baltic / Proto-Slavic

    Northern European hydronymy

    From Alteuropäische Hydronymie und urslavische Gewässernamen, by Udolph, Onomastica (1997), translated into English (emphasis mine):

    NOTE. An HTML version is available at Jurgen Udolph’s personal site.

    Because of the already striking similarities as the well-known “-m-case”, the number-words for ‘1000’, ’11’ and ’12’ and so on, J. Grimm had already assumed a close relationship between Germanic and Baltic and Slavic. (…)

    In my own search, I approached this trinity from the nomenclature side. In doing so, I noticed some name groups that can speak for a certain common context:

    1.* bhelgh-, *bholgh-.

    Map 10, p. 64, shows that a root * bhelgh- occurs in the name material of a region from which later Germanic, Baltic and Slavic originated. The Balkans play no role in this.

    bholgh-germanic-balto-slavic

    2. *dhelbh-, *dholbh-, *dhl̥bh-

    The proof of the three ablauting * dhelbh, * dholbh, * dhl̥bh- within a limited area shows the close relationship that this root has with the Indo-European basis. Again it is significant in which area the names meet (…)

    dhelbh-germanic-balto-slavic

    3. An Indo-European root extension *per-s- with the meaning ‘spray, splash, dust, drop’ is detectable in several languages (…). From a Baltic-Slavic-Germanic peculiarity cannot therefore be spoken from the toponymic point of view. The picture changes, however, if one includes the derived water names.

    4. The root extension *pel-t-, *pol-t-, *pl̥-t- of a tribe widely spread in the Indo-European languages around *pel-, pol- ‘pour, flow, etc.’, whose reflexes are found Armenian through Baltic and Slavic to the Celtic area, is found in the Baltic toponymy, cf. Latv. palts, palte ‘puddle, pool’.

    trzciniec-riesenbecher-culture
    The dynamics of stylistic changes of the form of the “Trzciniec pot” in the lowland regions of Central Europe, and spreading routes of the Trzciniec package in Central Europe. A good proxy for contacts through the Northern European Plain during the Early Bronze Age. Modified from Czebreszuk (1998).

    Early Balto-Finnic

    In order to properly delimit (geographically and chonologically) the Proto-Baltic and Proto-Slavic expansions, it is necessary to understand where the late Balto-Finnic homeland was located during the Bronze Age. The following are excerpts from the comprehensive hydrotoponymic study by Pauli Rahkonen (2013):

    In any case, Finnic probably had its origin somewhere around the Gulf of Finland. Names of large and central rivers such as Vuoksi (< Finnic vuo ‘stream’) and Neva (< Finnic neva ‘marsh, river’) must be very old and might represent Proto-Finnic hydronyms. In the southern coastal area of Finland, the names Kymi and Nietoo < *Niet|oja (id. later Porvoonjoki) may also be of Finnic origin and derive from, respectively, kymi ‘stream’ (see SSA I s.v. *kymi; see however SPK s.v. Kemijärvi; Rahkonen 2013: 24) and nieto(s) ‘heap of snow’ (SSA II s.v. nietos), in hydronyms probably ‘high (snowy?) banks of a river’. Mustion|joki is clearly a Finnish name < *must|oja ‘black river’. The river name Vantaa remains somewhat obscure, although Nissilä (see SPK s.v. Vantaanjoki) has derived it from the Finnic word vana ‘water route’. In western Finland the names of large rivers, such as Aura and Eura, are supposedly of Germanic origin (Koivulehto 1987).

    In Estonia the names of many of the most important rivers might be of Finnic origin: e.g. Ema|jõgi Est. ema ‘mother’ [Tartu district] (?? cf. the Lake Piiga|ndi < Est. piiga ‘maiden’), Pärnu [Pärnu district] < Est. pärn ‘linden’, Valge|jõgi [Loksa district] < Est. valge ‘white’, Must|jõgi [Võru district] < Est. must ‘black’. It is possible that Emajogi and especially Piigandi are the result of later folk etymologizing of a name with some unknown origin. However, as a naming motif there exist in Finland numerous toponyms with the stems Finnic *emä (e.g. 3 Emäjoki), *neit(V)- ‘maiden’ (e.g. Neitijärvi, Neittävänjoki, Neittävänjärvi) and Saami stems that can be derived from Proto Saami *nejte̮ ‘id’ (GT2000; NA).

    finnic-toponyms
    The historical southern boundary of Finnic hydronyms, excluding hydronyms produced by the Karelian refugees of the 17th century.

    These seemingly very old names of relatively large rivers in southern Finland, modern Leningrad oblast and Estonia support the hypothesis that Proto-Finnic was spoken for a long time on both sides of the Gulf of Finland and it thus basically corresponds to the hypothesis of Terho Itkonen (see below). In the Novgorod, Tver or Vologda oblasts of Russia, Finnic names for large rivers cannot be found (Rahkonen 2011: 229). For this reason, it is likely that the Late Proto-Finnic homeland was the area around the Gulf of Finland.

    Beyond the southeastern boundary of the modern or historically known Finnic-speaking area, there exists a toponymic layer belonging to the supposedly non-Finnic Novgorodian Čudes (see Rahkonen 2011). In theory it is possible that Proto-Finnic and Proto-Čudian separated from each other at an early stage or it is even possible that Proto-Čudian was identical with Proto-Finnic. However, this cannot be proven, because there is not enough material available describing what Novgorodian Čudic was like exactly.

    finno-saamic-mordvin
    Yakhr-, -khra, yedr-, -dra and yer-/yar, -er(o), -or(o) names of lakes in Central and North Russia and the possible boundary of the proto-language words *jäkra/ä and *järka/ä. Rahkonen (2013)

    A summary of the data is then:

    • The Daugava River and the Gulf of Livonia formed the most stable south-western Balto-Finnic border (up until ca. 1000 AD): the Daugava shows a likely Indo-European etymology, while some of its tributaries are best explained as derived from Uralic.
    • The first layer of “Early Baltic” loans in Early Balto-Finnic are of a non-attested Baltic dialect closest to Proto-Balto-Slavic (read more about this early layer).
    • The latest samples of the Trzciniec culture (or derived Iron Age group) from its easternmost group in Turlojiškė (ca. 1000-800 BC?) show a western shift towards Bell Beaker, although they show a majority of hg. R1a-Z280; while the earliest sample from Gustorzyn (ca. 1900 BC), likely from Trzciniec/Iwno, from the westernmost area of the culture, shows a Corded Ware-like ancestry (and hg. R1a-Z280, likely S24902+) among a BA sampling from Poland clearly derived from Bell Beaker groups.

    One can therefore infer that the expansion of the Trzciniec culture – as the earliest expansion of central-west European peoples into the Baltic after the Bell Beaker period – represented either the whole disintegrating Balto-Slavic community, or at least an Early Baltic-speaking community expanding from the West Baltic area to the east.

    The similarity of Early Slavs and the Trzciniec outlier with the Czech BA cluster, formed by samples from Bohemia (ca. 2200–1700 BC), and the varied haplogroups found among Early Slavs – reminiscent of the variability of the Unetice/Urnfield sampling – may help tentatively connect the early Proto-Slavic homeland more strongly with a Proto-Lusatian community immediately to the south-west of the Iwno/Proto-Trzciniec core.

    pca-late-bronze-age-balto-slavic-finnic
    Top Left:Likely Baltic, Slavic, and Balto-Finnic-speaking territories (asynchronous), overlaid over Late Bronze Age cultures. Balto-Slavic in green: West(-East?) Baltic (B1), unattested early Baltic (B2), and Slavic (S). Late Balto-Finnic (F) in cyan. In red, Tollense and Turlojiškė sampling. Dashed black line: Balto-Slavic/West Uralic hydrotoponymy border until ca. 1000 AD. Top right: PCA of groups from the Early Bronze Age to the Late Bronze Age. Marked are Iwno/Pre-Trzciniec of Gustorzyn (see below), Late Trzciniec/Iron Age samples from Turlojiškė, and in dashed line approximate extent of Tollense cluster; Y-DNA haplogroups during the Late Bronze Age (Bottom left) and during the Early Iron Age (Bottom right). Notice a majority non-R1a lineages among sampled Early Slavs. See full maps and PCAs.

    Proto-Balto-Slavic homeland

    Disconnected western border: Germanic

    The common Balto-Slavic – Germanic community must necessarily be traced back to the West Baltic. From Udolph’s Namenkundliche Studien zum Germanenproblem, de Gruyter (1994), translated from German (emphasis mine):

    My work [Namenkundliche Studien zum Germanenproblem] has shown how strong the Germanic toponymy is related to the East, less to Slavic, much more to Baltic. It confirms the recent thesis by W.P. Schmid on the special relationship Germanic and Baltic, according to which “the formation of the typical Germanic linguistic characteristics…must have taken place in the neighborhood of Baltic“.

    If one starts from a Germanic core area whose eastern boundary is to be set on the middle Elbe between the Erzgebirge and Altmark, there are little more than 400 km. to the undoubtedly Baltic settlement area east of the Vistula. Stretching the Baltic area westwards over the Vistula (as far as the much-cited Persante), the distance is reduced to less than 300 km. Assuming further that Indo-European tribes between the developing Germanic and the Baltic groups represent the connection between the two language groups, so can one understand well the special relationship proposed by W.P. Schmid between Germanic and Baltic. In an earlier period shared Slavic evidently the same similarities (Baltic-Slavic-Germanic peculiarities).

    balto-slavic-balto-finnic-homeland
    Top: Palaeo-Germanic (G2, blue area), Proto-Balto-Slavic/Pre-Baltic (PBSL, green area) and Early Proto-Balto-Finnic (PBF, cyan area) homelands superimposed over Early Bronze Age cultures. Persante hydronym and Gustorzyn ancient DNA sample location marked. Y-DNA haplogroups during the Early Bronze Age (Bottom left) and during the Middle Bronze Age (Bottom right). Notice a mix of R1b-L151 samples from the west and the process of integration of R1a-Z645 lineages from the the north-east. See full maps and PCAs.

    Substrate and immediate eastern border: Early Balto-Finnic

    While Balto-Finnic shows a late Balto-Slavic adstrate, Balto-Slavic has a Balto-Finnic(-like) substrate, also found later in Baltic and Slavic, which implies that Balto-Slavic (and later Baltic and Slavic) replaced the language of peoples who spoke Balto-Finnic(-like) languages, influencing at the same time the language of neighbouring peoples, who still spoke Balto-Finnic (or were directly connected to the Balto-Finnic community).

    For more on this relative chronology in Balto-Slavic – Balto-Finnic contacts, see e.g. the recent posts on Kallio (2003), Olander (2019), or a summary of this substrate.

    While Rahkonen (2013) entertains Parpola’s theory of a West-Uralic-speaking Netted Ware area (ca. 1900-500 BC), due to the Uralic-like hydrotoponymy of its territory, he also supports Itkonen’s idea of the ancient presence of almost exclusively Balto-Finnic place and river names in the Eastern Baltic and the Gulf of Finland since at least the Corded Ware period, due to the lack of Indo-European layers there:

    NOTE. This idea was also recently repeated by Kallio (2015), who can’t find a non-Uralic layer of hydrotoponymy in Balto-Finnic-speaking areas.

    It should be observed that the territory between the historical Finnic and Mordvin-speaking areas matches quite well with the area of the so-called Textile Ceramics [circa 1900–800 BC] (cf. Parpola 2012: 288). The culture of Textile Ceramics could function as a bridge between these two extreme points. Languages that were spoken later in this vast territory between Finland–Estonia and Mordovia seem to derive from Western Uralic (WU) as well. I have called those languages Meryan-Muroma, Eastern and Western Čudian and an unknown “x” language spoken in inland Finland, Karelia and the Lake Region of the Russian North (Rahkonen 2011; 241; 2012a: 19–27; 2013: 5– 43). This might mean that the territory of the Early Textile Ceramics reflects to some extent the area of late Western Uralic.

    The archaeologically problematic area is Estonia, Livonia and Coastal Finland – the area traditionally assumed to have been populated by the late Proto-Finns. The Textile Ceramics culture was absent there. It is very difficult to believe that the Textile Ware population in inland Finland migrated or was even the main factor bringing the Pre- or Early Proto-Finnic language to Estonia or Livonia. There are no archaeological or toponymic signs of it. Therefore, I am forced to believe that Textile Ceramics did not bring Uralic-speaking people to those regions. This makes it possible, but not absolutely proven, to assume that some type of Uralic language was spoken in the region of the Gulf of Finland already before Textile Ceramics spread to the northwest (circa 1900 BC).

    corded-ware-west-uralic
    Top Left: Corded Ware culture expansion. Top right: PCA of Corded Ware and Sub-Neolithic groups. Y-DNA haplogroups during the Corded Ware expansion (Bottom left) and during the subsequent Bell Beaker expansion (Bottom right). Notice the rapid population replacement of typical Corded Ware R1a-Z645 lineages by expanding Bell Beakers of hg. R1b-L23 in central-east Europe, while they show continuity in the described ancestral Fennoscandian West-Uralic-speaking territory. See full maps and PCAs.

    The Corded Ware population in Finland is thought to have been NW Indo-European by many scholars (e.g. Koivulehto 2006: 154–155; Carpelan & Parpola 2001: 84). At least, it is probable that the Corded Ware culture was brought to Finland by waves of migration, because the representatives of the former Late Comb Ceramics partially lived at the same time side by side with the Corded Ware population. However, it is possible that the immigrants were a population that spoke Proto-Uralic, who had adopted the Corded Ware culture from their Indo-European neighbors, possibly from the population of the Fatjanovo culture, e.g. in the Valdai region. This was suggested by Terho Itkonen (1997: 251) as well. In that case the population of the Typical and Late Comb Ceramics may have spoken some Paleo European language (see Saarikivi 2004a). In the Early Bronze Age, the Baltic Pre-Finnic language that I have suggested must have been very close to late WU and therefore no substantial linguistic differences existed between the Baltic Pre-Finns and the population of Textile Ceramics in inland Finland. I admit that this model is difficult to prove, but I have presented it primarily in order to offer new models of thinking.16 At least, there is no archaeological or linguistic reason against this idea.

    This dubitative attribution of Proto-Uralic to the expansion of Corded Ware groups in eastern Europe, which is what hydrotoponymic data suggests in combination with archaeology, has to be understood as a consequence of how striking Rahkonen finds the results of his research, despite Itkonen’s previous proposal, in the context of an overwhelming majority of Indo-Europeanists who, until very recently, simplistically associated Corded Ware with the Indo-European expansion.

    Conclusion

    Even Kortlandt accepts at this point the identification of expanding East Bell Beakers from the Carpathian Basin as those who left the Alteuropäische layer reaching up to the Baltic. However, he identified Udolph’s data solely with West Indo-European, forgetting to mention the commonly agreed upon western Proto-Balto-Slavic homeland, most likely because it contradicts two of his main tenets:

    1. that Balto-Slavic split from a hypothetical Indo-Slavonic (i.e. Satem) group expanding from the east; and
    2. that laryngeals can be reconstructed for Balto-Slavic – unlike for North-West Indo-European.
    old-european-asian-hydro-toponymy
    Indo-European hydrotoponymy in Europe and the Middle East (scarce Central Asian data). Baltic data compensated, statistical method RBF: intermediate regions devoid of Indo-European toponyms are inferred to have them; it compensates thus e.g. for the scarce Indo-European hydrotoponyms in Poland by assuming ‘soft’ continuity from West Germany to the Baltic.

    A hypothetic “Pre-Indo-Slavonic” laryngeal Indo-European layer reaching Fennoscandia and the Forest Zone with Corded Ware is fully at odds with all known data:

    • in comparative grammar, since the one feature that characterizes Graeco-Aryan is precisely its set of innovations relative to Northern Indo-European, which presupposes a longer contact (and further laryngeal loss) once Tocharian and North-West Indo-European had separated – hence probably represented by Palaeo-BalkanCatacomb-Poltavka contacts once Afanasevo and Yamna settlers from the Carpathian Basin / East Bell Beakers had become isolated;
    • in hydrotoponymy, because of the prehistoric linguistic areas that can be inferred from (1) the distribution of Old European hydrotoponymy; (2) Udolph’s work on Germanic and the likely non-Indo-European substrate in Scandinavia and land contacts with Balto-Finnic; (3) from the Northern European traits in the Northern European Plain; or (4) from the decreasing proportion of Indo-European place and river names from central Europe towards the east and north.
    • NOTE. An alternative explanation of Old European/Balto-Slavic layers, e.g. by a ‘Centum’ Temematic – even if one obviates the general academic rejection to Holzer’s proposal – couldn’t account for the absolute lack of an ancestral layer of Indo-European hydrotoponymy in North-Eastern Europe (i.e. the longest-lasting Corded Ware territory), in sharp contrast with Western Europe, South-Eastern Europe, and South Asia. All of that contradicts an Eastern Indo-European community, even without a need to recall that the oldest hydrotoponymic layers common to Fennoscandia and the Forest Zone are of Uralic nature.

    • in archaeology, because cultural expansions of the Eastern European Early Bronze Age province since the Bell Beaker period (viz. Mierzanowice, Trzciniec, Lusatian, Pomeranian, West Baltic Culture of Cairns) suggest once and again west-east movements, most (if not all) of which – based on the presence of Indo-European speakers during the common era – were likely associated with Indo-European-speaking communities replacing or displacing previous ones.
    • in palaeogenomics, because of the late and different association of Corded Ware ancestry and haplogroups among Balto-Slavic and Indo-Iranian communities, in turn corresponding to the different satemization processes found in both dialects, which may have actually been related to the Uralic substrate that is found in both (read more on Uralic influences on Balto-Slavic and on Indo-Iranian).

    On the other hand, a careful combination of Uralic and Indo-European comparative grammar, hydrotoponymic data, and population genomics fits perfectly well Itkonen’s and Rahkonen’s association of Corded Ware in Eastern Europe with Uralic languages, as well as the traditional mainstream view of Uralic before Indo-European in Fennoscandia and in the Forest Zone, as I explained in a recent post about genetic continuity in the East Baltic area.

    Population genomics is not the main reason to reject the Indo-European Corded Ware theory – or any other prehistoric ethnolinguistic identification, for that matter. It can’t be. This new field offers just the occasional confirmation of a well-founded theory or, alternatively, another nail in the coffin of fringe theories that were actually never that likely, but seemed impossible to fully dismiss on purely theoretical grounds.

    The problem with Corded Ware was that we couldn’t see how unlikely its association with Indo-European languages was until we had ancient DNA to corroborate archaeological models, because few (if any) Indo-Europeanists really cared about the linguistic prehistory of eastern and northern Europe, or about Uralic languages in general (contrary to the general trend among Uralicists to be well-versed in Indo-European studies). Now they will.

    Related

    European hydrotoponymy (III): from Old European to Palaeo-Germanic and the Nordwestblock

    nordic-bronze-age-cultures

    The study of hydrotoponymy shows a prevalent initial Old European layer in central and northern Germany, too, similar to the case in Iberia, France, Italy, and the British Isles.

    The recent paper on Late Proto-Indo-European migrations by Frederik Kortlandt relies precisely on this ancestral layer as described by Jürgen Udolph to support a Danubian expansion of North-West Indo-European with East Bell Beakers, identified as the Alteuropäische (Old European) layer that was succeeded by Germanic in the North European Plain.

    The Proto-Germanic homeland

    The following are excerpts are translated from the German original (emphasis mine) in Udolph’s Namenkundliche Studien zum Germanenproblem, de Gruyter (1994):

    udolph-namenkunde
    Buy the book at De Gruyter’s site or at Amazon.

    The following is a concise compilation of the investigation into nine points, which will be subsequently discussed: there are Brink (in the north brekk-), -by (on the Elbe), the name of the Elbe itself, germ, haugaz and blaiw, klint, malm / melm, the name of the Rhön, and the place name element -wedel.

    I want to briefly summarize the results:

    1. Brink has toponymically a clear focus in Germany between the Rhine and the Weser; in Schleswig-Holstein and Denmark it is almost completely missing, the Scandinavian place name documents show an accumulation in eastern Sweden. The English Brink names can not be associated with the Scandinavian ones. The “real” Scandinavian variant brekka, brekke, however, also appear on the Shetland and Orkney Islands and in central England.

    2. The Central Elbian –by-place names have nothing to do with the Danish and Scandinavian -by-names.

    3. The name of the Elbe has been carried from south to north and has become an appellative in Scandinavia. This clearly proves that a south-north migration has taken place.

    4. The distribution of haugaz does not support a Nordic origin of the word. K. Bischoff in his thorough investigation never asked whether the reverse path from south to north would be possible. However, in comparison with the results of the study of other toponyms, this second option will be much more likely to be accepted. On the “problem of the gap” in the distribution (between Aller and northern Holstein) see page 910.

    hlaiwaz-germanisch

    5. Completely missing is the assumption of Nordic origin in the case of hlaiwaz. A look at Map 67 shows this clearly.

    6. Even in the case of klint, Denmark and Scandinavia are only marginally involved in the distribution of names. This contradicts the thesis that the English Klint names are of Nordic origin. On the other hand, Map 68 (Klit- / Klett-) shows how Nordic place names can have an influence on the British Isles.

    klint-germanisch

    7. Even in the case of germ, melm (ablauting malm, mulm), everything speaks for a continental Germanic starting point: here are all ablaut stages in the appellative vocabulary and in the toponymy, which shows together with the name Melmer perhaps the most ancient -r-derivations, which are unknown to the Nordic area, while the Nordic names, in turn, have a distinct tendency to spread to eastern Sweden, towards the Baltic Sea.

    8. The name of the Rhön can only be interpreted with the aid of the Nord Germanic apellative hraun “boulder field, stony ground, lava field”. This does not mean that Nord Germanic peoples have given this name, but that the Common or Proto-Germanic peoples knew the appelative still. The Rhön owes its name to this language stage.

    9. The spread of the fronds names in Germany, classified by E. Schröder as “North Germanic invasion”, can be explained differently: more important than the often younger names north of the Elbe in Schleswig-Holstein (type Wedelboek) are the place names near Braunschweig, Büren (Westphalia), and in the Netherlands, in which case a south-north spread is more convincing than the assumption of a Nordic expansion.

    wedel-germanisch

    If you take the similar distribution maps 15 (wik), 31 (fenn), 36 (slk), 39 (büttel), 47 (live), 49 (quem), 50 (thing), 61 (brink) and 66 (haugaz) It can be seen from this (page 72, page 908) that there are parts of Germany which, to a lesser degree, are more heavily involved than others in Old Germanic place name formations: that applies to southern Thuringia, the Area between Werra and Fulda, the Magdeburger Börde and its western foothills to the Weser at the Porta Westfalica). On the other hand, the areas north of the Aller, Hanoverian Wendland and wide areas between the Lower Weser and the Lower Elbe (apart from the area around Osterholz-Scharmbeck as well as Kehdingen and Hadeln) are little and hardly affected.

    There is no question that the reasons for the different dispersion can not lie in the name itself, but have other causes. H. Kuhn has considered the natural conditions of the landscape with the fronds. Comparing the place name expansion outlined here with a bog map of Lower Saxony, as found in numerous publications (Map 73, page 910), solves the problems: even today’s bog distribution of Lower Saxony, diminished through cultivation and drainage (albeit still considerable), reflects the fact that the early colonization and naming of northern Germany has been shaped and, to a certain extent, controlled by settler-friendly and not-settler-friendly conditions.

    moorkarte-deutschland
    Distribution of bogs in Germany. Source: M. Sommer, Institut für Bodenlandschaftsforschung, ZALF, Müncheberg.

    On the location of the Germanic Urheimat

    According to the space briefly outlined by the present study, the Old Germanic settlement area in toponymic terms is roughly to be located between the Erzgebirge, Thüringerwald, Elbe, Aller and an open border in Westphalia, for the following reasons:

    • High proportion of old European names. This is a basic requirement, which of course is also fulfilled by other areas, but not by Schleswig-Holstein, Denmark and Scandinavia. (…)
    • Of particular importance was the discussion about relations with the north (the generally accepted ancient Germanic settlement area, section L, p. 830-917). I believe that the detailed study of the geographical names no longer allows one to assume a Scandinavian homeland of Germanic tribes. Too many arguments speak against it. It is much more likely to start with a northward migration (…).
    bell-beaker-germanic
    Bell Beaker expansion ca. 2600-2200 BC. Top Left: Tentative location of the Pre-Proto-Germanic homeland (earliest stage), in the North European Plain between the Elbe and the the Aller (open border). Top right: PCA of the Bell Beaker period, with Netherlands EBA cluster (population west of the Germanic Urheimat) in red, and Battle Axe/Baltic CWC (population east and north of the Urheimat) in cyan. Bottom left: ADMIXTURE analysis of ancient DNA samples. Bottom right: Y-DNA haplogroup map. See full maps and PCAs.

    Western border: Nordwestblock

    Recently, W. Meid has once more dealt in detail with Kuhn’s thesis. After that, the most important criteria for the approach of this thesis are the following:

    1. -p- (and other shutter sounds) are partly not shifted in North German names;
    2. the existence of a -sí-suffix;
    3. -apa in river names;
    4. the suffix -andr-;
    5. certain words u. Name strains, e.g. Veneter, Belgian.
    6. Above-average relations of the northwestern block to Italic (Latin, Osco-Umbrian).

    W. Meid agrees with Kuhn’s theses, but with limitations: “These evidences seem to indicate that the NW-space did not belong to the original settlement area of ​​the Teutons, but that the Germanization of this area or larger parts of it did not take place until relatively late, namely – as Kuhn thinks – after the Germanic sound shift or during its last phase. According to Kuhn’s own words this “space… appears as a block that has long defied Germanization”.

    Udolph continues explaining why most of these non-Germanic examples are “optic illusions”, since he can explain most of them as from Old European to Old Germanic stages, which is mostly in agreement with the known features of Old European hydrotoponymy. For example, -apa- and -andra-names as Old European; -p- as before the Germanic sound shift; -st- and -s-formations as Northern European; -ithi- also unrelated to a hypothetic “Venetic” substrate.

    I think that the point to discuss should not be the similarity with Old European or the oldest reconstructible Proto-Germanic stage (i.e. the closest to North-West Indo-European), or the appearance of these traits also in neighbouring Germanic territory, but the proportion of “more archaic” features contrasting with the proper Germanic area, and thus differences in frequency with the Germanic core territories.

    Just as Udolph can’t accept the non-Indo-European nature of most cases, one can’t simply accept his preference for a Pre-Proto-Germanic nature either, for the same reason one can’t accept the relationship of Western European “Pre-Celtic” hydrotoponymy with Celtic peoples because of some shared appellatives whose Celtic nature is not proven.

    NOTE. If there is something missing from this huge book is certainly statistical analyses with GIS, which would make this case much easier to discuss in graphical and numerical terms. Let’s hope Udolph can update the data in the near future, because he is still (fortunately) active.

    In any case, the Nordwestblock remains a likely Old European hydrotoponymic area partially shared by Germanic, which doesn’t lie at the core of the spread of Old European place names and has a potential non-Indo-European substrate shared with Northern European groups. Combined with comparative grammar and with results of population genomics supporting the spread of East Bell Beakers of Yamna descent from the Carpathian Basin, this essentially renders interpretations of Old European expansion from Northern Europe devoid of support in linguistics.

    Palaeo-Germanic expansion

    To the north, the settlement movement depends on the location and spread of settlement-deficient areas, such as the moors northeast of Wolfsburg, north of Gifhorn, south of Fallingbostel, etc. As soon as this belt has been breached, the place name frequency in the eastern Lüneburg Heath indicates where more favorable settlement conditions are to be found: the Altmark in Saxony-Anhalt, the Jeetzel lowlands and especially the Ilmenau area near Uelzen, Bevensen and Lüneburg (it is difficult not to recall the name Jastorf here).

    If one combines these findings with the dispersion of ancient Germanic place names, one will find that above all the section of the river east from Hamburg to about Lauenburg was particularly favorable for crossing. The onomastic data speaks in favour of this aspect, e.g. the following names lying north and south of this area.

    brink-germanisch

    1. Delvenau = Elbe-Lübeck Canal.

    2. Neetze north of Lüneburg (-d-/-t-change).

    3. Wipperau north of Lüneburg (-p-/-b- change).

    4. The dispersion of the -wik places (Bardowik), cf. Map 15, p. 106.

    5. The dissemination of the -r formations (Map 24, p. 191).

    6. The -ithi formations Geesthacht, Bleckede u.a. south of the Elbe, Eckede north of the stream (see Map 28, p.272).

    7. Fenn south of the Elbe in the north of Lüneburg (Map 31, p.315).

    8. The distribution of the Hor name (Harburg) and northeast of it in Holstein (Map 32, p.328).

    9. Germ, sik- with clear clusters southeast. and northeastern. from Hamburg (Map 36, p. 409).

    10. Also the -büttel names show a concentration east of Hamburg on the one hand and a second accumulation at the estuary of the Elbe (Brunsbüttel) (map 39, p.438).

    11. Gorleben and other places in Hann. Wendland south of the river (Map 47, p.503).

    12. Werber-names southeast from Hamburg and in eastern Holstein (Map 53, p.742).

    13. The scattering of brink names (Map 61, p. 843).

    The place name distributions also make it possible to track the settlement movement north of the Elbe. It has been repeatedly emphasized that Schleswig-Holstein has little share in old Germanic toponymy. One tries to explain this fact, which reaches into the realm of the Old European hydronyms, by saying that, according to archeology, “large parts of Schleswig-Holstein in the 5th to 7th centuries were sparsely populated”.

    scandinavia-neolithic-dagger-period
    Close contacts in Fennoscandia. The distribution of Scandinavian flint daggers (A) in the east and south Baltic region and possible trends of “down the line” trade (B). Good size and quality flint zone in the south-west Baltic region is hatched (C). According to: Wojciechowski 1976; Olausson 1983, fig. 1; Madsen 1993, 126; Libera 2001; Kriiska & Tvauri 2002, 86. Image modified from Piličiauskas (2010).

    If one summarizes these synoptically (Map 74, p.914) and also takes into account the not-included -leben-names (Map 47, p.503), then it is quite clear that Denmark by no means shares these types of names. The most important points are, in my opinion:

    1. North of today’s German-Danish border, the quantity of old place names drops rapidly and even tends towards zero. West Jutland in particular is rarely involved in the dispersion.
    2. Within Jutland there is a clear orientation to the east. The connection with southern Sweden is established via Funen and Zeeland.
    3. Disputed is in my opinion, whether the spread of toponymy followed a roughly direct line Fehmarn and Lolland/Falster. This is not to be excluded, but the maps of toponymy distribution do not give a clear indication in this direction.

    The synoptic map makes it clear that both western Schleswig-Holstein and western Jutland are not to be regarded as Old Germanic settlement areas. Rather, East Jutland and the Danish islands were reached by Germanic tribes.

    pca-bronze-age-germanic
    Bronze Age groups ca. 2200-1750 BC. Top Left: Tentative location of (1) the Pre-Proto-Germanic homeland (earliest stage), in the North European Plain between the Elbe and the the Aller (open border), (2) the Pre-Proto-Germanic expansion area, coinciding with the Nordic Dagger Period, and (3) the Pre-Proto-Germanic-like Nord-West-Block. Top right: PCA of European Bronze Age groups. Bottom left: ADMIXTURE analysis of ancient DNA samples. Bottom right: Y-DNA haplogroup map. See full maps and PCAs.

    Absolute chronology and Balto-Finnic

    It is imprecise to estimate the age of settlement movements from toponymic research. I do not want to be involved in speculation, but I think that Klingberg’s estimate could have some arguments in its favor. In the approximate dating, however, it is important to include a fact that has already been briefly mentioned above and should be treated here in more detail: the fact of Germanic-Finnic relations.

    W.P. Schmid has emphatically pointed out the difficulty that arises when one considers the unfolding of Germanic too far from the Baltic Sea settlement areas. Among other things, it draws attention to the fact that a Germanic homeland that were postulated too far west could not explain how Germanic loanwords might appear in the Finnic names of Northern Russia. These will be mentioned with reference to M. Vasmer: Randale to Finn. ranta “beach”, Pel’doza and Nimpel’da to Finn. pelto, Justozero to Finn. juusto “cheese”, Tervozero to Finn. terva “tar” and Rovdina Gora to Finn. rauta “ore”.

    I think it is possible that the clear spread of Old and North Germanic toponyms, as described in the synoptic map 74 (p. 914) and in the already mentioned -ing, -lösa, -by, -sta(d) and -säter-maps (19, 46, 63-65), can offer some help: quite early the Germanic tribes reached the Swedish east coast. It is also clear that there have previously been contacts with Slavic and Finno-Ugric tribes by sea. However, intensive German-Finnic relations can, in my opinion, have come about only through close contacts on the mainland.

    Pre-Indo-European substrate

    In my investigation, I have repeatedly come up with suggestions to explain a hard-to-interpret North Germanic name from a Pre-Germanic, possibly Non-Indo-European substrate. Most of these were views of H. Kuhn, which he also used to support his so-called “Nord-West block”.

    On one point H. Kuhn may have been right with an assumption of a Pre-Germanic substrate that did not provide the basis for further development in Germanic terms: he very clearly argued that Scandinavia too was Pre-Germanic, even Pre-Indo-European A substrate that stands out above all because of the lack of Lautverschiebung : “In the Nordic countries, we have to reckon with non-Germanic, non-Indo-European prehistoric names scarcely less than in the other Germanic languages”. In light of the results of the present work that makes a relatively late Germanization of Scandinavia very likely, this sentence should not be set aside in the future, but carefully examined on the basis of the material.

    Both data, the known long-lasting Palaeo-Germanic – Finno-Samic contacts, and the underresearched presence of non-Indo-European vocabulary in Scandinavia, are likely related to the presence of a West Uralic(-like) substrate in Scandinavia and most likely also in Northern Europe, based on the disputed non-Indo-European components shared through the North European Plain (see above), and on the scarce ancient Indo-European hydrotoponymy in central-east Europe to the north of the Carpathians.

    Population genomics

    Although there is yet scarce genetic data from northern European territories, the haplogroup distribution among sampled peoples from the Germanic migration period and during the Viking expansion suggests a prevalence of R1b-U106 in the North European Plain (also found in Barbed Wire Beakers), and thus a later integration of typically Neolithic (I1) and CWC-related (R1a) subclades to the Germanic-speaking community during the expansion into Southern Scandinavia.

    This is compatible with the described development of maritime elites by Bell Beakers, representing maritime mobility and trade, and an appealing ideology, similar to the prevalence of Athens over Sparta (Corded Ware in this analogy). It is also supported by the bottlenecks under R1b-U106 to the north of Schleswig-Holstein.

    NOTE. Nevertheless, other R1b-L151 may have been part of the Germanic-speaking communities, especially during its earliest stage, and also R1b-U106 (and other R1b-L161) subclades may appear all the way from the Carpathians to Northern Europe, including the Eastern European Early Bronze Age.

    germanic-iron-age
    Common Germanic expansions ca. 500 BC on. Top Left: Early Iron Age cultures. Top right: PCA of groups from the Iron Age to the Middle Ages. Y-DNA haplogroups during the Germanic migrations (Bottom left) and during the Middle Ages (Bottom right). Notice a majority of R1b-U106 (practically absent from previous Bronze Age populations of Central Europe) among sampled Germanic tribes. See full maps and PCAs.

    Archaeology

    This sudden population bust to the south and predominance of a Southern Scandinavian maritime society in the Nordic circle seems to be also supported by inferences from archaeological data, too. For example, from the recent Human impact and population dynamics in the Neolithic and Bronze Age: Multi-proxy evidence from north-western Central Europe, by Feeser et al. The Holocene (2019):

    The second boom between c. 3000 and 2900 cal. BC relates to increases in the palynological proxy and the binned all site SCDPD curve. From an archaeological point of view, this time reflects the transition from the Funnelbeaker to the Single Grave Culture. The emergence of this new cultural phenomenon is often regarded to have been associated with a shift in subsistence practices, that is, a shift from sedentary agricultural to mobile pastoral subsistence (Hinz, 2015; Hübner, 2005; Iversen, 2013; Sangmeister, 1972).

    denmark-demography-bronze-age
    Left: Map with pollen sites. Right: Bin sensitivity plots based on summed calibrated date probability distributions (SPD) using different degrees of binning on-site level (h = 0 no binning; h = 1000 high binning) and Kernel density plots (KDE) of available radiocarbon dates from the settlement context (settlement sites). Modified from the paper to include a red arrow showing Corded Ware bust and subsequent boom with the Dagger Period..

    (…) there is palynological evidence for increased importance of cereal cultivation during the Young Neolithic in comparison to the Early Neolithic (Feeser et al., 2012). This, however, does not rule out an increased importance of pastoralism, as grazing on grasslands and extensive cereal cultivation are difficult to distinguish and to disentangle in the palynological record. Generally however, human impact on the environment and population levels, respectively, did not reach Funnelbeaker times maxima values during this boom phase at the beginning of the Younger Neolithic. The similar short-term synchronous developments in both the pollen profiles during 2800–2300 cal. BC could point to large-scale, over-regional uniform development during the Younger Neolithic in our study area (cf. also Feeser et al., 2016).

    Between c. 2400 and 2300 cal. BC, the palynological proxy and the binned all site SCDPD curve show a similar distinct decrease (Figure 6), and we define a second bust phase accordingly. The soil erosion record, however, indicates elevated values at around this time but declines, although not very well defined, to a minimum at around 2200 cal. BC. Due to the generally low number of colluvial deposits recorded for the Younger Neolithic, this is not regarded to contradict our interpretation, as low sample sizes generally minimize the chances of identifying a robust pattern. A strong increase in all the three proxies between 2200 and 2100 cal. BC defines our third boom phase.

    Bronze Age evolution

    Candidate homelands for the succeeding (Palaeo-Germanic) stages of the language are shifted also in archaeology to the south, due to the economic influence of demographically stronger Nordic Bronze Age cultural groups of northern Germany over Southern Scandinavia.

    A good description of societal changes in the Palaeo-Germanic stages is offered by the recent paper Cultural change and population dynamics during the Bronze Age: Integrating archaeological and palaeoenvironmental evidence for Schleswig-Holstein, Northern Germany, by Kneisel et al. The Holocene (2019):

    schleswig-holstein-culture-demography
    Qualitative data from material culture and demography in Schleswig-Holstein and Mecklenburg-Western Pomerania. Modified from the original to remark periods of likely demographic decrease (red square) and growth (blue square).

    At each beginning of a boom phase and each end of a bust phase, changes in the material culture could be observed.

    When the pressure on the landscape is at its lowest around 1500 BC and shortly before it rises again, the type of burial changes, hoards and bronzes increase, and monumental burial mounds are erected again. Vice versa, when the pressure on the landscape reaches its maximum value around 1250 BC, tools and hoard depositions decrease again and only the monumental burial and prestige goods are maintained. The ‘elite’ are continuing with their way of burial. The reduction in house surface area and the number of hoards takes place earlier, possibly because of material scarcity as could also be proven in Thy, northern Jutland (Bech and Rasmussen 2018).

    Again, the human impact decreases, and at its lowest point at the beginning of Period IV ca. 1100 BC, the monumental burial custom and the addition of prestige goods also end. The number of hoards and graves begins to rise again, and cooking pits appear. Exchange networks shift with the beginning of Period V, while axes increase again together with a slight decrease in the human impact curve. The appearance of certain artefacts or burial rites at the beginning of such a period of upheaval seems to suggest the role of a trigger. With this analysis, we have defined several likely indicators for social change in the less distinct phases and societal change in the strongly pronounced phases around 1500 BC and 1100 BC and the most important triggers for the Schleswig-Holstein Bronze Age.

    soegel-wohlde-nordic-bronze-age
    Distribution of burials with Valsømagle, Sögel and Wohlde blades with provenance known to parish. q = Valsømagle blades; s = Wohlde blades (small = one grave with a blade; medium = two graves with a blade); l = Sögel blades (small = one grave with a blade, medium = two graves with a blade, large = three graves with a blade). From Bergerbrant (2007).

    While population movements can’t be really understood without a proper genetic transect proving or disproving archaeological theories, it seems that the intermediate zone of the Nordic circle was subjected to at least two demographic busts and succeeding booms during the Middle and Late Bronze Age periods, which not only affected the hydrotoponymy of Schleswig-Holstein (see above), but probably served as dynamic changes in the linguistic evolution of Palaeo-Germanic-speaking communities up to the Common Germanic expansion.

    Read more on the Northern Early Bronze Age province.

    Related

    Volosovo hunter-gatherers started to disappear earlier than previously believed

    volosovo-corded-ware

    Recent paper (behind paywall) Marmot incisors and bear tooth pendants in Volosovo hunter-gatherer burials. New radiocarbon and stable isotope data from the Sakhtysh complex, Upper-Volga region, by Macānea, Nordqvist, and Kostyleva, J. Archaeol. Sci. (2019) 26:101908.

    Interesting excerpts (emphasis mine):

    The Sakhtysh micro-region is located in the Volga-Oka interfluve, along the headwaters of the Koyka River in the Ivanovo Region, central European Russia (Fig. 1). The area has evidence of human habitation from the Early Mesolithic to the Iron Age, and includes altogether 11 long-term and seasonal settlements (Sakhtysh I–II, IIa, III–IV, VII–XI, XIV) and four artefact scatters (sites V–VI, XII–XIII), in addition to which burials have been detected at five sites (I–II, IIa, VII, VIII) (Kostyleva and Utkin, 2010). The locations have been known since the 1930s and intensively studied since the 1960s under the leadership of D.A. Kraynov, M.G. Zhilin, E.L. Kostyleva, and A.V. Utkin.

    Sakhtysh II and IIa are the most extensively studied sites of the complex, with ca. 1500m2 and around 800m2 excavated, respectively. The burial grounds at both sites are considered as fully investigated.

    volosovo-sakhtysh-dates
    AMS datings from the sites Sakhtysh II and IIa. Sampled contexts are given in parentheses (burial/hoard), “crust” indicates samples of charred organic
    residues on pottery from cultural layer. For data, see Tables 1–2.

    Sakhtysh chronology

    The AMS dates do not support the previously proposed phasing of the Sakhtysh burials to early (4750–4375 BP/3600–3000 cal BCE), late (or developed; 4375–4000 BP/3000–2500 cal BCE), and final (4000–3750 BP/2500–2200 cal BCE): the early and late burials at Sakhtysh IIa do not stand out as two separate groups, and also the burials and hoards from Sakhtysh II, connected to the final phase, are temporally overlapping with these. Neither the use sequence, where the settlement and burial phases are non-overlapping and also complementary between the sites (Kostyleva and Utkin, 2010, 2014), finds support in the present material.

    The AMS datings indicate that the Volosovo people started to bury their dead at Sakhtysh IIa after 3700 cal BCE; dates earlier than this may be affected by FRE or suffer from mixed contexts and poor quality of dates. The present data questions the interpretation that the Sakhtysh IIa cemetery was used without interruptions between 4800 and 4080 BP (Kostyleva and Utkin, 2010), i.e. for a millennium between 3550 and 2600 cal BCE. The AMS dates rather suggest a use period of some centuries only around the mid-4th millennium cal BCE, tentatively 3650–3400 cal BCE. This would also be more realistic considering the number of burials at the site.

    volosovo-sakhtysh-sites
    The core area of Volosovo culture (after Kraynov, 1987) and the sites of the Sakhtysh complex (after Kostyleva and Utkin, 2010). Eurasian map base made with Natural Earth. Illustration: K. Nordqvist.

    Volosovo chronology

    The absolute dating of Volosovo culture was for a long time hampered by the small number of radiocarbon dates (see Kraynov, 1987). Today,>100 datings connected with it can be found in literature (Korolev and Shalapinin, 2010; Chernykh et al., 2011; Nikitin, 2012; Mosin et al., 2014). Unfortunately, the available dates do not form solid grounds for dating the cultural phenomenon, as many of them have quality-related issues, large measurement errors, and ambiguous cultural or physical contexts. Consequently, particular datings may be connected to different cultural phases by different scholars. Finally, a large part of the newly-published datings are obtained through direct dating of potsherds (Kovaliukh and Skripkin, 2007; Zaitseva et al., 2009), and therefore, their cogency must be faced with reservation (see Van der Plicht et al., 2016; Dolbunova et al., 2017).

    The datings connected with Volosovo cover a wide time range between ca. 5500 BP (4400 cal BCE) and ca. 3700 BP (2100 cal BCE). However, datings from secure contexts, with good quality (error ca. 50 years or below) and no probable FRE, place the beginning of Volosovo culture to the first half of the 4th millennium cal BCE, around 3700–3600 cal BCE. This is also supported by the roughly coeval terminal dates given for the preceding Lyalovo (Zaretskaya and Kostyleva, 2011) and Volga-Kama cultures (Lychagina, 2018), as well as the appearance of related neighbouring cultures, for example, in the Kama region (Nikitin, 2012; Lychagina, 2018), the southern forest steppe area (Korolev and Shalapinin, 2014), and north-western Russia and Finland (Nordqvist, 2018). Still, the dating of many of these cultural phases suffers from the same problems as of Volosovo.

    A handful of contested datings place the end of Volosovo culture to the final centuries of the 3rd millennium cal BCE, or even later (Kostyleva and Utkin, 2010; Chernykh et al., 2011; Nikitin, 2012). On the other hand, the new AMS dates indicate that Volosovo activities at Sakhtysh II and IIa ceased before or towards the early 3rd millennium cal BCE; if this reflects the general decline of Volosovo culture must be still confirmed by more dates from Sakhtysh and elsewhere. In this context, the general cultural development must be accounted for. To what extent – if at all – the Volosovo people were present after the arrival of the Corded Ware culture-related Fatyanovo-Balanovo populations? Based on the current, albeit scant and inconclusive radiocarbon data this took place from ca. 2700 cal BCE onwards (Krenke et al., 2013).

    volosovo-fatyanovo-balanovo
    Corded Ware and Comb Ware hunter-gatherer-related populations in north-eastern Europe from ca. 2600 BC. See full map.

    Comments

    One of the interesting genetic papers in the near future will be the one that finally includes samples from Corded Ware groups in the forest zone (i.e. Fatyanovo-Balanovo and Abashevo), which will most likely confirm that they are the origin of the known genetic profile of Central and East Uralic-speaking peoples, seeing how West Uralic peoples show genetic continuity in the East Baltic area, coinciding with the Battle Axe culture.

    Uralicists have come a long way from the 1990s, when the picture of Uralic before Balto-Slavic in the Baltic was already evident, and Uralians were identified with Comb Ware peoples. The linguistic data and relative chronology are still valid, despite the now outdated interpretations of absolute archaeological chronology, as happens with interpretations of Krahe or Villar about Old European.

    As an example, here are some relevant excerpts from Languages in the Prehistoric Baltic Sea Region, by Kallio (2003):

    NOTE. Kallio’s contribution appeared in the book Languages in Prehistoric Europe (2003), which I hold nostalgically close in my Indo-European library (now almost impossible to read fully). It is still one of my preferred books (from those made up of mostly unconnected chunks on European linguistic prehistory), because it contains Oettinger’s essential update of North-West Indo-European common vocabulary, which led us indirectly to our Modern Indo-European project from 2005 on.

    In any case, the Uralic arrival in the region east of the Baltic Sea preceded the Indo-European one (…).

    This theory that the ancestors of Finno-Saamic speakers arrived in the Baltic Sea region earlier than those of Balto-Slavic speakers is still rejected by some scholars (e.g. Napolskikh 1993: 41-44), who claim, for instance, that Finno-Saamic speakers would not have known salmons before they met Balts because the Finno-Saamic word for ‘salmon’ (i.e. *losi) is a borrowing from Baltic. Similarly, one could claim that English speakers would not have known salmons before they met Frenchmen because English salmon is a borrowing from French. In other words, Worter und Sachen are not necessarily borrowed hand in hand. Otherwise, it would not be so easy to explain how many Finnish names of body parts are borrowings from Baltic (e.g. hammas ‘tooth’, kaula ‘neck’, reisi ‘thigh’) and from Germanic (e.g. hartia ‘shoulder’, lantio ‘loin’, maha ‘stomach’).

    A more probative argument is the fact that Balto-Slavic features in Finno-Saamic are mostly lexical ones (i.e. typical superstrate features), where Finno-Saamic features in Balto-Slavic are mostly non-lexical ones (i.e. typical substrate features). Note that there are more Balto-Slavic features in Finnic than in Saamic and more Finno-Saamic features in Baltic than in Slavic. This fact could be explained by presuming that Pre-Saamic was spoken north of the Corded Ware area and Pre-Slavic was spoken south of the Typical Pit-Comb Ware area, whereas Pre-Finnic and Pre-Baltic alone were spoken in the area, where both the Typical Pit-Comb Ware culture (ca. 4000-3600 BC) and the Corded Ware culture (ca. 3200-2300 BC) were situated. This area was most probably bilingual, until Finnic and Baltic won in the north and in the south, respectively.

    As is well-known, the idea of Uralic substrate features in Balto-Slavic is not new (cf. e.g. Pokorny 1936/1968: 181-185). As recent studies (e.g. Bednarczuk 1997) have shown, their density is the most remarkable in the four Balto-Slavic languages spoken in the earlier Pit-Comb Ware area (i.e. Latvian, Lithuanian, Belorussian, Russian). On the other hand, occasional Uralisms in the other Balto-Slavic languages spoken west of the Vistula and south of the Pripyat may rather be considered adstrate features spread from the northeast.

    comb-ware-uralic
    Our beliefs from the 2000s. A hypothetic Uralic Comb Ware distribution before the arrival of a hypothetic North-West Indo-European-speaking Corded Ware. “Generalized distribution of the Pit-Comb Ware cultural complex (Mallory & Adams 1997: 430, Carpelan 1999: 257) and the most probable homelands of Saamic, Finnic, Mordvin, Mari, and Permic.”

    The idea of Indo-European superstrate features in Finnic is not new either (cf. e.g. Posti 1953). As Jorma Koivulehto (1983) has recently shown, the earliest Indo-European loanword stratum in the westernmost Uralic branches alone can be considered Northwest Indo-European and connected with the Corded Ware culture (ca. 3200-2300 BC). Since this layer, there have been continuous contacts between Baltic and Finnic. According to Koivulehto (1990), the following stratum can be called Proto-Balt(o-Slav)ic and dated to the Late Neolithic period (ca. 2300-1500 BC). Note that this Proto-Balt(o-Slav)ic dating agrees with the established ones (cf. e.g. Shevelov 1964: 613-614, Kortlandt 1982: 181), when we remember the fact that archaeologists have also moved their datings back by centuries during the last decades.

    Finally, there is also a Baltic loanword stratum which was not borrowed from the ancestral stage of Latvian, Lithuanian and/or Old Prussian but from some extinct Baltic language or dialect (Nieminen 1957). However, as these words still go back to the early Proto-Finnic stage, they can hardly be dated later than Bronze-Age ( ca. 1500-500 BC). Therefore, we may conclude that they were probably borrowed from a Baltic superstrate, which arrived in the Finnish Gulf area during the Corded Ware period and survived there until the Bronze Age, when it was no longer identical with other Baltic dialects. In any case, as later Baltic loanword strata concern southern Finnic languages alone, we may presume that this ‘North Baltic’superstrate had become extinct.

    The traditional association of Uralic with Volosovo hunter-gatherers doesn’t make sense, since they neither miraculously survived for thousands of years nor mixed for hundreds of years with Corded Ware peoples, so we can now more confidently reject the recent assumption by Carpelan & Parpola that their language was adopted by incoming Fatyanovo, Balanovo and Abashevo groups, to develop into the known Uralic languages (more here). This includes one of the many models of the the Copenhagen group, who simplistically follow “Steppe ancestry” for Indo-Europeannes.

    If one combines the known relative linguistic chronology with the North-West Indo-European hydrotoponymy layer, now more clearly identified as Old Europeans expanding with East Bell Beakers and derived Early Bronze Age groups, I think there is little space left for maneuvering out of the overwhelming evidence for a Uralic homeland in the forest-steppes, linked to the spread of late Sredni Stog/Corded Ware ancestry into north-eastern Europe and beyond the Urals.

    Related

    A Song of Sheep and Horses, revised edition, now available as printed books

    cover-song-sheep-and-horses

    As I said 6 months ago, 2019 is a tough year to write a blog, because this was going to be a complex regional election year and therefore a time of political promises, hence tenure offers too. Now the preliminary offers have been made, elections have passed, but the timing has slightly shifted toward 2020. So I may have the time, but not really any benefit of dedicating too much effort to the blog, and a lot of potential benefit of dedicating any time to evaluable scientific work.

    On the other hand, I saw some potential benefit for publishing texts with ISBNs, hence the updates to the text and the preparation of these printed copies of the books, just in case. While Spain’s accreditation agency has some hard rules for becoming a tenured professor, especially for medical associates (whose years of professional experience are almost worthless compared to published peer-reviewed papers), it is quite flexible in assessing one’s merits.

    However, regional and/or autonomous entities are not, and need an official identifier and preferably printed versions to evaluate publications, such as an ISBN for books. I took thus some time about a month ago to update the texts and supplementary materials, to publish a printed copy of the books with Amazon. The first copies have arrived, and they look good.

    series-song-sheep-horses-cover

    Corrections and Additions

    Titles
    I have changed the names and order of the books, as I intended for the first publication – as some of you may have noticed when the linguistic book was referred to as the third volume in some parts. In the first concept I just wanted to emphasize that the linguistic work had priority over the rest. Now the whole series and the linguistic volume don’t share the same name, and I hope this added clarity is for the better, despite the linguistic volume being the third one.

    Uralic dialects
    I have changed the nomenclature for Uralic dialects, as I said recently. I haven’t really modified anything deeper than that, because – unlike adding new information from population genomics – this would require for me to do a thorough research of the most recent publications of Uralic comparative grammar, and I just can’t begin with that right now.

    Anyway, the use of terms like Finno-Ugric or Finno-Samic is as correct now for the reconstructed forms as it was before the change in nomenclature.

    west-east-uralic-schema

    Mediterranean
    The most interesting recent genetic data has come from Iberia and the Mediterranean. Lacking direct data from the Italian Peninsula (and thus from the emergence of the Etruscan and Rhaetian ethnolinguistic community), it is becoming clearer how some quite early waves of Indo-Europeans and non-Indo-Europeans expanded and shrank – at least in West Iberia, West Mediterranean, and France.

    Finno-Ugric
    Some of the main updates to the text have been made to the sections on Finno-Ugric populations, because some interesting new genetic data (especially Y-DNA) have been published in the past months. This is especially true for Baltic Finns and for Ugric populations.

    ananino-culture-new

    Balto-Slavic
    Consequently, and somehow unsurprisingly, the Balto-Slavic section has been affected by this; e.g. by the identification of Early Slavs likely with central-eastern populations dominated by (at least some subclades of) hg. I2a-L621 and E1b-V13.

    Maps
    I have updated some cultural borders in the prehistoric maps, and the maps with Y-DNA and mtDNA. I have also added one new version of the Early Bronze age map, to better reflect the most likely location of Indo-European languages in the Early European Bronze Age.

    As those in software programming will understand, major changes in the files that are used for maps and graphics come with an increasing risk of additional errors, so I would not be surprised if some major ones would be found (I already spotted three of them). Feel free to communicate these errors in any way you see fit.

    bronze-age-early-indo-european
    European Early Bronze Age: tentative langage map based on linguistics, archaeology, and genetics.

    SNPs
    I have selected more conservative SNPs in certain controversial cases.

    I have also deleted most SNP-related footnotes and replaced them with the marking of each individual tentative SNP, leaving only those footnotes that give important specific information, because:

    • My way of referencing tentative SNP authors did not make it clear which samples were tentative, if there were more than one.
    • It was probably not necessary to see four names repeated 100 times over.
    • Often I don’t really know if the person I have listed as author of the SNP call is the true author – unless I saw the full SNP data posted directly – or just someone who reposted the results.
    • Sometimes there are more than one author of SNPs for a certain sample, but I might have added just one for all.
    ancient-dna-all
    More than 6000 ancient DNA samples compiled to date.

    For a centralized file to host the names of those responsible for the unofficial/tentative SNPs used in the text – and to correct them if necessary -, readers will be eventually able to use Phylogeographer‘s tool for ancient Y-DNA, for which they use (partly) the same data I compiled, adding Y-Full‘s nomenclature and references. You can see another map tool in ArcGIS.

    NOTE. As I say in the text, if the final working map tool does not deliver the names, I will publish another supplementary table to the text, listing all tentative SNPs with their respective author(s).

    If you are interested in ancient Y-DNA and you want to help develop comprehensive and precise maps of ancient Y-DNA and mtDNA haplogroups, you can contact Hunter Provyn at Phylogeographer.com. You can also find more about phylogeography projects at Iain McDonald’s website.

    Graphics
    I have also added more samples to both the “Asian” and the “European” PCAs, and to the ADMIXTURE analyses, too.

    I previously used certain samples prepared by amateurs from BAM files (like Botai, Okunevo, or Hittites), and the results were obviously less than satisfactory – hence my criticism of the lack of publication of prepared files by the most famous labs, especially the Copenhagen group.

    Fortunately for all of us, most published datasets are free, so we don’t have to reinvent the wheel. I criticized genetic labs for not releasing all data, so now it is time for praise, at least for one of them: thank you to all responsible at the Reich Lab for this great merged dataset, which includes samples from other labs.

    NOTE. I would like to make my tiny contribution here, for beginners interested in working with these files, so I will update – whenever I have time – the “How To” sections of this blog for PCAs, PCA3d, and ADMIXTURE.

    -iron-age-europe-romans
    Detail of the PCA of European Iron Age populations. See full versions.

    ADMIXTURE
    For unsupervised ADMIXTURE in the maps, a K=5 is selected based on the CV, giving a kind of visual WHG : NWAN : CHG/IN : EHG : ENA, but with Steppe ancestry “in between”. Higher K gave worse CV, which I guess depends on the many ancient and modern samples selected (and on the fact that many samples are repeated from different sources in my files, because I did not have time to filter them all individually).

    I found some interesting component shared by Central European populations in K=7 to K=9 (from CEU Bell Beakers to Denmark LN to Hungarian EBA to Iberia BA, in a sort of “CEU BBC ancestry” potentially related to North-West Indo-Europeans), but still, I prefer to go for a theoretically more correct visualization instead of cherry-picking the ‘best-looking’ results.

    Since I made fun of the search for “Siberian ancestry” in coloured components in Tambets et al. 2018, I have to be consistent and preferred to avoid doing the same here…

    qpAdm
    In the first publication (in January) and subsequent minor revisions until March, I trusted analyses and ancestry estimates reported by amateurs in 2018, which I used for the text adding my own interpretations. Most of them have been refuted in papers from 2019, as you probably know if you have followed this blog (see very recent examples here, here, or here), compelling me to delete or change them again, and again, and again. I don’t have experience from previous years, although the current pattern must have been evidently repeated many times over, or else we would be still talking about such previous analyses as being confirmed today…

    I wanted to be one step ahead of peer-reviewed publications in the books, but I prefer now to go for something safe in the book series, rather than having one potentially interesting prediction – which may or may not be right – and ten huge mistakes that I would have helped to endlessly redistribute among my readers (online and now in print) based on some cherry-picked pairwise comparisons. This is especially true when predictions of “Steppe“- and/or “Siberian“-related ancestry have been published, which, for some reason, seem to go horribly wrong most of the time.

    I am sure whole books can be written about why and how this happened (and how this is going to keep happening), based on psychology and sociology, but the reasons are irrelevant, and that would be a futile effort; like writing books about glottochronology and its intermittent popularity due to misunderstood scientist trends. The most efficient way to deal with this problem is to avoid such information altogether, because – as you can see in the current revised text – they wouldn’t really add anything essential to the content of these books, anyway.

    Continue reading

    Official site of the book series:
    A Song of Sheep and Horses: eurafrasia nostratica, eurasia indouralica

    Genetic continuity among Uralic-speaking cultures in north-eastern Europe

    east-europe-bronze-age

    The recent study of Estonian Late Bronze Age/Iron Age samples has shown, as expected, large genetic continuity of Corded Ware populations in the East Baltic area, where West Uralic is known to have been spoken since at least the Early Bronze Age.

    The most interesting news was that, unexpectedly for many, the impact of “Siberian ancestry” (whatever that actually means) was small, slow, and gradual, with slight increases found up to the Middle Ages, compatible with multiple contact events in north-eastern Europe. Haplogroup N became prevalent among Finnic populations only through late bottlenecks, as research of modern populations have long suggested, and as ancient DNA research hinted since at least 2015.

    I risked to correlate the arrival of chiefs from the south-west with the infiltration of N1c-VL29 subclades during the transition to the Iron Age, coupled with that minimal “Siberian” ancestry (see e.g. here and here). Now we know that the penetration of this non-CW ancestry started, as predicted, in the Iron Age; that it was highly variable in the few samples where it appeared, with ca. 1-4%, while most Iron Age individuals show 0%; and that it was not especially linked to individuals of N1c-Vl29 lineages.

    It is also basically confirmed, based on the (ancient and Modern Swedish) N1c-L550 subclades found among Iron Age Estonians, that N1c-VL29 lineages and the so-called “Siberian” ancestry will be found simultaneously around the Baltic coastal areas, and that different lineages must have suffered later founder effects among Finns, which suggests that these alliances through exogamy brought exactly as much language change in Sweden, Lithuania, or Poland, as they did in the East Baltic region…

    On the other hand, the paper has also shown a potential movement of Corded Ware-derived peoples, if the change from LBA to IA samples is meaningful; in fact, even more Corded Ware-like than Baltic and Estonian BA populations. The exact origin of that movement is difficult to pinpoint, and it may not be related to the arrival of Akozino warrior-traders from the south-east, since theirs seems to be a minor impact proper of elites in a chiefdom system around the Baltic.

    fortified-settlements-lba-ia
    Distribution of fortified settlements (filled circles) and other hilltop sites (empty circles) of the Late Bronze Age and Pre-Roman Iron Ages in the East Baltic region. Tentative area of most intensive contacts between Baltic and Balto-Finnic communities marked with a dashed line. Image modified from (Lang 2016).

    Also suggesting a potential movement is the ‘southern’ shift observed in the West and East Baltic areas, likely showing the arrival of Proto-East Baltic speakers (such as the Trzciniec outlier), as we have already discussed in this blog. The unexpected increase in Corded Ware-like ancestry in the Eastern Baltic, coupled with the expected large continuity of hg. R1a-Z283 in the homeland of Balto-Finnic expansions, gives even more support to the known complex system of exogamy along the Baltic coasts, and offers another potential reason for the rise of Baltic-speaking territories in the West Baltic: elite domination.

    It is nevertheless important to understand that, even among the most “genetic continuous” regions like Estonia, not a single population in Europe is heir of some ancestral, immutable people. Not in terms of haplogroups, and not in terms of admixture. Balto-Finnic speakers, however continuous they might seem (e.g. in Southern Estonians) aren’t an exception.

    After all, this blog was (re)born to fight the currently prevalent sheer stupidity surrounding the simplistic “R1a/steppe ancestry=Indo-European” association, so I wouldn’t like to see it replaced with some other stupid continuity or purity ideas within 10 to 20 years…

    Late Uralic stems from East Corded Ware groups

    With the currently available tools – linguistics, archaeology, and now genetics -, I don’t think there is any argument to date to question the direct connection of the Late Proto-Uralic expansion with all Eastern Corded Ware groups (i.e. Battle Axe, Fatyanovo-Balanovo, and Abashevo), and thus at least with the unifying A-horizon of Corded Ware and the bottlenecks under R1a-Z645.

    NOTE. The only out-group among Corded Ware cultures is the Single Grave culture. It appears to be an early Corded Ware offshoot, reflected in their non-unitary cultural traits (distinct from later unifying waves), in their varied patrilineal clans, and in the short-lasting cultural effect in northern Europe before their complete demise under pressure of expanding Yamna/Bell Beaker peoples from the Danube. The culture’s minimal (if any) effects on succeeding peoples might be seen mostly in the (mainly phonetic) Uralic substrate found in Balto-Slavic – although this may also stem from a more eastern influence, close to the Baltic – and in the contacts of Celtic with Uralic. The huge time depth between this early hypothetic Uralic layer in northern Europe and the emergence of peoples inhabiting these territories in recorded history have no doubt been erroneously interpreted as a lack of Uralic presence in the area.

    1) That connection was evident in the Yamna – CWC differences in archaeology, and especially later, with at least Fatyanovo-Balanovo and Abashevo representing the obvious replacement of the Volosovo culture before further expansions of CWC-related groups west and east of the Urals.

    The mythical millennia-long continuity of Volosovo hunter-gatherers, including centuries among Corded Ware peoples, as expected lately by the Copenhagen group (and anyone who doesn’t want to question the 1960s association of Indo-European with CWC) must be rejected today in population genomics, as the recent studies of ancient and modern populations show, and as ancient DNA from the region will confirm.

    2) In linguistics, the survival of Volosovo as The Uralic-speaking culture was also hardly believable. From Kallio (2015):

    While we can say at least something about Uralic substrates in Northeastern Europe, non-Uralic substrates cannot at all easily be identified, because of multiple language shifts, viz. first from non-Uralic to Uralic and then from Uralic to Russian. Yet the Soviet Uralicist Boris Serebrennikov (1956, 1959) argued that there are some non-Uralic substrate toponyms in the Volga-Oka region, but his idea was never taken seriously in the west (cf. Sauvageot 1958), and it pretty soon also sank into oblivion in Russia, even though it can still occasionally pop up there in non-onomastic circles (cf. Napolskikh 1995: 18–19). However, not all the hypotheses on non-Uralic substrates in Northeastern Europe should be rejected (see e.g. Helimski 2001b).

    bronze-age-early-languages-east-europe
    Tentative map of the distribution of known languages in Eastern Europe during the Early Bronze Age. See full map.

    Helimski (2001) argues for a non-Uralic topo-hydronomy in Northern Russia, whose population may have kept their languages up to the Common Era despite the Corded Ware expansion, which is in line with the survival of some non-Indo-European languages everywhere in Europe after the expansion of Yamna and its offshoots:

    It should be borne in mind that these [Uralic] hydronyms reached us mainly through Northern Russian and, accordingly, with a tendency to phonetic-morphological adaptation and unification (for river names it is “natural” to be, like the word ‘river’ itself, feminine and to end in -a). Taking into account this circumstance, it may turn out to be non-useless for etymological identification of at least some of the hydronyms on the Finno-Ugric basis.

    On the other hand, I wouldn’t exclude the possibility that some parts of this large geographical area were never (completely) Finno-Ugric. The population that created the most important part of the hydronymy of the Russian North could be finally pushed aside or assimilated only at the end of the 1st – beginning of the 2nd millennium AD, during the Russian colonization, retaining the memory of the White-Eyed Chude in its own memory.

    NOTE. For more on this non-IE substrate in (especially West) Uralic, see e.g. Zhivlov (2015),

    The same non-Uralic substrate is most likely behind most of the shared traits by Mordvinic and Balto-Finnic (see below).

    3) In genetics, I don’t think the picture could get any clearer. I don’t know what “Steppe ancestry = Indo-European” proponents expected from 2019, if they expected anything at all (I haven’t seen any coherent model, proposal, or prediction for a long time now), but I doubt the recent results are compatible with any of their implied expectations.

    corded-ware-pca-sub-neolithic-europe
    Detail of the PCA of the Corded Ware expansion. See full PCA and more related files.

    Notice, from the PCA above, how this Baltic Late Neolithic group shows actually a shift from Sredni Stog (see PCA with Sredni Stog) towards typical Khvalynsk-Urals-related ancestry, i.e. populations from eastern European forested regions, derived from hunter-gatherer pottery groups, as I have proposed for a very long time, since the first time a Baltic LN “outlier” appeared. It’s amazing how some amateurs can find 0.1% of any Siberian outlier’s ancestry among Uralians 4,000 years later, but fail to see the direct connection here. The esoteric uses of qpAdm, I guess…

    Especially noticeable is the extra WHG-like ancestry and corresponding shift, seen especially marked in late Polish CWC samples, but also in Baltic CWC and especially in one Sweden Battle Axe sample, all of them shifting apparently closer to Pitted Ware and SHG. While that may have been interpreted as an in situ admixture in Scandinavia before, the late Polish CWC samples show likely a resurgence of local populations, so we can assume that both shifts (to SHG- and EHG-like populations) of available CWC samples around the Baltic are clearly part of the WHG:EHG continuum that will be found in the eastern European sub-Neolithic cultures, from Narva to Volosovo.

    This WHG-related ancestry is clearly predominant in groups with which Battle Axe peoples admixed, based on the shift towards Pitted Ware, which – I can only guess based on modern Volga Finns – is different from the shift we will see in Netted Ware, more towards the Khvalynsk-Urals cluster. This is in line with the expansion of Battle Axe eastward through coastal areas (West to East Baltic and Finland into Sweden), while Fatyanovo peoples probably emerged from a slightly different route, but also a northern one, if one is to follow archaological similarities and their chronology.

    bronze-age-europe-baltic
    Detail of the PCA of European Bronze Age populations. See full PCA and more related files.

    During the Iron Age, the only peoples that probably shifted strongly (based on modern populations) are West Baltic ones, getting closer to the available Late Trzciniec samples, and even closer to the Trzciniec outlier, i.e. away from the earlier Eastern Corded Ware cluster, and towards Central European groups like Czech EBA or Poland EBA, both of them clearly derived from Bell Beakers, but also admixed with (and thus shifted toward) CW-like populations.

    If one looks carefully at the previous PCA on Bronze Age populations, and the next one on Iron Age clusters, it is evident that adding the Swedish LN outlier to East Baltic BA (both strongly related to Battle Axe populations) essentially gives us the continuity of East Baltic BA into the Iron Age. This cluster is continued also in two outliers from Sigtuna, a Viking town close to the Gulf of Finland, known to be an important trading site, 1,500 years later. Not much of a change around the Gulf of Finland, then:

    iron-age-eastern-europe
    Detail of the PCA of East and North European Iron Age populations. See full PCA and more related files.

    Based on the two simplistic Uralic clines one might see described (among the many that certainly existed, from Corded Ware to different Eurasian populations), and just like BOO was for some months fashionable as “Samic”, some may be tempted to say that certain Sintashta or Srubna outliers close to the Urals mark the True Uralic™ peoples. Because, of course they do. Ghost haplogroup N and stuff. And Corded Ware never ever Uralic. Because Gimbutas, and my IE R1a grandfather.

    NOTE. Funny thing here: there might be Corded Ware, Iranian, Slavic, Germanic, etc… outliers or out-groups, and they might form the widest genetic clusters ever seen, but they are all of one language, because archaeology and linguistics; however, one “outlier” (also, put your own definition of “outlier” here, let’s say 1% of whatever, and strontium isotope potentially from 100 km away) ca. 600 BC in the Baltic who (surprise!) happens to show hg. N, and he signals the first incoming True Uralic™ speaker from wherever… It won’t be the first or the last time some people resort to “the complexity of Uralic-speaking peoples” in ancestry, just to look for “hg. N = Uralic” like crazy. You only need common sense to understand that this is not how this works. Amateur genomics can’t get more embarrassing than the current “let’s look for ‘Siberian ancestry’ in every individual of haplogroup N” trend. Or maybe it can, and it will, but I can’t see it yet.

    If one were to insist on looking for ‘foreign’ contributions among Iron Age Estonians, though, I think one should also check out first archaeology, and then the PC3 (or, more graphically, a 3D plot), to understand what might be happening with the many Uralic clines derived from Corded Ware, before starting to play around with bioinformatic tools to discover a teeny tiny 1% admixture of the wrong population, and rushing to build far-fetched narratives. Apparently, one of the different clines formed roughly between southern (steppe – forest-steppe) and northern (tundra-taiga) populations in Uralians is also seen in some Iron Age Estonian individuals – especially in some late samples from Ingria…This is not my main interest, so I will leave this here for others to keep wasting their time chasing the white whale of the 0.5% of True Uralic™ ancestry in ancient Baltic samples of hg. N.

    pca-3d-estonians-iron-age-boo-samic
    Still images of the 3D plot of Eurasian samples. Typical PC1 vs. PC2 visualization to the left, and shift of the view to PC3 on the right image. See full PCA and more related files.

    An exclusive Volga-Kama homeland for Disintegrating Uralic?

    Since I don’t believe in macro-regions of largely continuous ethnolinguistic communities, as I have often said about Slavic (naively associated with prehistoric tribes of Eastern Europe) or Germanic (absurdly considered to be represented by Battle Axe), it is difficult for me to believe that Battle Axe-derived cultures remained of the same Finno-Samic dialects since the Corded Ware expansion…unless we live in Westeros, where everything happens “for thousands of years”.

    I have to admit, then, that the now prevalent identification among Uralicists has become quite attractive:

    • Fatyanovo-Balanovo as Finno-Permic:
      • Fatyanovo/Netted Ware with West Uralic (also called Finno-Mordvinic).
      • Balanovo/Chirkovo-Kazan with Central Uralic (Mari-Permic).
    • Abashevo, into the Andronovo-like Horizon through the Seima-Turbino phenomenon, with East Uralic (also Ugro-Samoyedic).

    Exactly like the identification of Yamna Hungary – Bell Beaker transition as the North-West Indo-European homeland, it gives us simplicity and small and late ethnolinguistic communities, away from the traditionally overused big and early language territories.

    This late homeland would be supported, among others, by:

    • The presence of Indo-Iranian loanwords in Finno-Permic and Ugric (probably also in Samoyedic, either lost, or – much more likely – underresearched), compatible with the immediate contact between Abashevo – Sintashta-Potapovka-Filatovka and Fatyanovo-Balanovo.
    • The supposed expansion of Netted Ware from Fatyanovo to the north-west, which may be explained as the split and expansion of Balto-Finnic and Samic ca. 1900 BC.
    • A longer-lasting Finno-Permic (West+Central Uralic) community contrasting with the early separation of East Uralic.
    • The compatibility of this late expansion with the late expansion of Pre-Germanic from Denmark with the Dagger Period, and of Balto-Slavic with Trzciniec, which puts all three dialects reaching the Baltic Sea in the EBA.

    NOTE. I meant to update the linguistic text to include the most recently favoured phylogenetic tree of Uralic languages after Häkkinen (2007, 2009, 2014), which has very quickly become the new normal among Uralicists, but I don’t think I will have enough time to review the necessary papers for that. I am rushing to publish a printed edition, so the text will wind up being a mixture of “traditional” (meaning, basically, pre-2010s) description of Uralic dialects but using modern divisions; say, “West Uralic” instead of “Finno-Samic”. By the way, I am still amazed that none of my reader-haters (or any online user discussing Uralic migrations, for that matter) have come up with the questions that the new division pose, and it supports my suspicion about the complete lack of interest in linguistics of most (a)DNA fans, except for the occasional use of old and free PDFs Googled to support new narratives invented expressly for some qpAdm results…

    textile-ceramics-europe-bronze-age
    Textile ceramic styles and influence of Bronze Age cultures divided in clusters.

    Problems with this Parpola-Carpelan’s (2012-2018) interpretation include:

    • The differentiation between Fennoscandian Textile Ceramics vs. Netted Ware, which is not warranted in archaeology. The assumption that Netted Ware expanded to the Baltic Sea (as Kallio does, following the traditional view) is thus weak, and it was probably a question of cultural contacts coupled with short-distance population movements/exchange in both directions (from the Baltic to the Volga and vice versa). In fact, the culture division relies on some fairly common and technically simple ornamentation patterns, widespread all over northern Europe, even before the Corded Ware expansion, and it is very difficult to separate certain neighboring Textile Ceramics from Netted Ware groups in southern Finland (i.e. Sarsa-Tomitsa groups).
    • The strict and radical direction described for the Netted Ware by Carpelan, as an eastward and northward expansion, within a very short time frame (ca. 1900-1800 BC), based on few radiocarbon dates, which seems to me like a very risky assumption. We know how this kind of descriptions of direction of culture expansion based on radiocarbon dates has turned out in much more complex “packages”, like the Bell Beaker culture… In fact, the earliest dates for Textile Ware are from the East Baltic, earlier than those of Netted Ware.
    • The assumption that Balto-Finnic traits shared with Mordvinic are a) late and b) meaningful for dialectalization of two closely related dialects, when it is clear that both dialects separated quite early. Phonologically Finnic is more conservative, morphologically less so, and the shared traits include a handful of non-Uralic substrate words which can’t be traced to a single common source, hence they were adopted when both languages had already separated… All in all, Finnic – Mordvinic correspondances are not even close to Italo-Celtic ones, which is clearly fully incompatible with a proposal of a Finnic separation from Mordvinic coinciding with the LBA-IA transition.

    Especially problematic for Parpola’s model is the lack of genetic impact in Bronze Age or Iron Age Estonians, not reaching a significant level under any possible statistical threshold – which I am sure was quite disappointing for some of my readers -, but is in line with major archaeological continuity of groups the from region, only disturbed in cultural (and Y-chromosome) terms by the expansion of Akozino warrior-traders all over the Baltic Sea. Any proposed population movement will be very difficult to support in genetics, given the Corded Ware-derived populations that we will see in both regions, and the continued Baltic-Volga contacts since the Corded Ware expansion.

    Problems with an interpretation of such a small impact in population genomics includes the similarly weak impacts and haplogroup infiltrations that can be seen among populations basically everywhere in Eurasia, during any given period, and much greater genetic impacts that are supposed to be (or that were certainly) followed by ethnolinguistic continuity.

    akozino-malar-axes-fennoscandia
    Distribution of the Akozino-Mälar axes according to Sergej V. Kuz’minykh (1996: 8, Abb. 2).

    The Battle Axe question

    From Kallio (2015), about choosing a tentative homeland for Proto-Uralic:

    (…) linguistically uniform Proto-Uralic would have been spoken in the Volga-Oka region until the mid-third millennium BC when the Proto-Uralic-speaking area would have expanded to the Volga-Kama region as well. By the end of the same millennium, this expansion would have led to the earliest dialectal splits within Uralic into Finno-Mordvin, Mari-Permic, and Ugro-Samoyed. The splitting up of these three soon followed during the early second millennium BC when the Uralic-speaking area finally stretched from the Baltic Sea in the west to the Altai mountains in the east. Indeed, no matter where Proto-Uralic was spoken, the branching into the nine well-attested subgroups (viz. Finnic, Saami, Mordvin, Mari, Permic, Hungarian, Mansi, Khanty, and Samoyed) must have taken less than a millennium, because their shared phonological and morphosyntactic isoglosses are rather limited (see Salminen 2002). The traditional view that all this branching would have taken several millennia violates everything linguistic typology teaches us about the rate of language change.

    The basic problem of this identification of Fatyanovo-Balanovo as West-Central Uralic and Abashevo as East Uralic is the nature of the Battle Axe culture, including the Bronze Age East Baltic and Gulf of Finland area. Even if it is accepted that Fatyanovo-Balanovo represented all Western groups, Battle Axe must have represented West Uralic-like dialects.

    The ethnolinguistic identification of Battle Axe depends ultimately on the nature of contacts of Fatyanovo/Netted Ware with Battle Axe/Textile Ceramics. If both groups were close and interacted profusely, as it seems, it doesn’t seem granted that we will be able to distinguish a close Para-West Uralic dialect of Scandinavia from the actual expanding Balto-Finnic and Samic dialects, if they were actually linked to the Netted Ware expansion. Also from Kallio (2015):

    No doubt the most convincing substrate theory has recently been put forward by the Saami Uralicist Ante Aikio (2004), who has not only rehabilitated but also improved the old idea of a non-Uralic substrate in Saami. His study shows that there were still non-Uralic languages spoken in Northern Fennoscandia as recently as the first millennium AD. Most of all, they were not only genetically non-Uralic but also typologically non-Uralic-looking, bearing a closer resemblance to the so-called Palaeo-European substrates (for which see e.g. Schrijver 2001; Vennemann 2003).

    In comparison, the case of Finnic is much more difficult. The fact that Proto-Uralic was not spoken in the East Baltic region means that this area must have originally been non-Uralic-speaking, but so far the evidence for a non-Uralic substrate in Finnic has consisted of appellatives and proper names with no etymology (cf. Ariste 1971; Saarikivi 2004a). Contrary to the proposed substrate words in Saami, those in Finnic show no structural non-Uralisms, as if they had indeed been borrowed from some genetically related or at least typologically similar languages, as I suggested above. Also none of them is more recent than the Middle Proto-Finnic stage, which makes them at least two millennia old. All this agrees with archaeological evidence discussed earlier that the Uralicization of the East Baltic region occurred during the Bronze Age (ca. 1900–500 BC).

    The discussion of the paper continues with an unsuccessful attempt to find a hypothetical ancient Indo-European substrate that Kallio believes must be associated with the expansion of Corded Ware, in line with the traditional belief. For example, the often mentioned – almost folk etymology-like, unsurprisingly popular among amateurs – ‘Neva’ as derived from IE “young” is logically rejected…Unlike Parpola, Kallio’s view seems to be confident that Netted Ware (as Textile Ware) expanded into the East Baltic, on both sides of the Gulf of Finland, already during the Bronze Age.

    As it has become apparent in population genomics, none of them was right, and Textile Ceramics will essentially show – like Netted Ware – a large genetic continuity of Corded Ware peoples in the whole north-eastern European forest zone – despite small regional population movements, obviously -, which necessarily implies that the whole Corded Ware culture – and not only Fatyanovo-Balanovo and Abashevo – were Uralic-speaking territories.

    The similarities in terms of culture and Y-DNA bottlenecks between Battle Axe and Fatyanovo-Balanovo also imply that the linguistic differences between these groups were probably not many, and became strongly divided only after their territorial division. Continued contacts between Battle Axe- and Fatyanovo-derived groups can explain the proposed contacts (Finnic with Samic, Finnic with Mordvinic) after their linguistic-but-not-physical separation.

    east-european-fatyanovocwc
    East European movement directions (arrows) of the representatives of the Central European Corded Ware Culture (according to I.I. Artemenko).

    Battle Axe spoke “Para-Balto-Finnic”?

    The Balto-Finnic-speaking nature of Battle Axe is thus supported by:

    • The lack of non-Uralic substrates in Balto-Finnic territory (Kallio 2015).
    • The early separation of Samic and Finnic from Mordvinic, and the virtual identity of Proto-West-Uralic and Proto-Uralic, which suggests that Proto-Uralic spread fast (Parpola 2012).
    • The scarce non-Uralic topo-hydronymy in the East Baltic and around the Gulf of Finland (Saarikivi 2004), comparable to that on the Upper Volga region.
    • The strong influence of a Balto-Finnic-like substrate on Pre-Germanic (or, in Kallio’s opinion, the same Scandinavian substrate influencing both Germanic and Balto-Finnic at the same time), and the continued influence of Balto-Finnic on Proto-Baltic and Proto-Slavic.
    • The continued influence of Corded Ware-derived groups in central-east Sweden in Finland and the East Baltic in terms of agricultural innovations appearing in the LBA, compatible with Schrijver’s proposal of intermediate Germanic-shifted Balto-Finnic groups and Balto-Finnic groups influenced by their pronunciation.
    • The intense Palaeo-Germanic and late Balto-Slavic / early Proto-Baltic superstrate on Balto-Finnic, which place all three dialects around the Baltic Sea since the Early Bronze Age.
    • The easy replacement of a hypothetic Para-Balto-Finnic dialect by incoming Proto-Balto-Finnic-speaking peoples (say, with textile ceramics), without much linguistic impact.

    In fact, the continuous contacts of the East Baltic with the Volga, and especially the close interaction with Akozino warrior-traders just before the Tarand-grave period, could be the actual origin of the recent (if any) Finnic-Mordvinic connections that need to be traced back to the LBA-IA (maybe here the number ‘ten’), since most of them can be related to a Pit-Comb Ware culture substrate and earlier contacts through the forest zone, which Samic (due to its early split and presence to the north of the Gulf of Finland during the BA) does not share. In fact, some of them can be traced back to Balto-Finnic first

    These are the most often mentioned, in order of descending relevance for a shared ancient community:

    • Noun paradigms and the form and function of individual cases.
    • The geminate *mm (foreign to Proto-Uralic before the development of Fennic under Germanic influence) and other non-Uralic consonant clusters.
    • The change of numeral *luka ‘ten’ with (non-Uralic) *kümmen.
    • The presence of loanwords of non-Uralic origin, related to farming and trees, potentially Palaeo-European in nature.

    It’s not only a question of quantity. Are these shared Mordvinic – Balto-Finnic traits really more relevant than, say, those between Italo-Celtic, which are supposed to have formed a community for a very short period at the end of the 3rd millennium around the Alps? Are these traits even sufficient to propose a common early Mordvinic-Finnic group within West Uralic, rather than loose Mordvinic – Balto-Finnic contacts, i.e. contacts between East Baltic (Textile Ceramics) and Volga-Kama (Netted Ware)?

    Based on the alternative (Kallio’s) view of continued contacts between Textile Ceramics groups, even without knowing anything about linguistics, you can guess that Parpola is spinning very thin when assuming that these changes suggest that Balto-Finnic may have expanded with Akozino warrior-traders, separating thus ca. 800 BC from Mordvinic…

    Genetic findings now clearly help dismiss any meaningful population impact in the LBA-IA transition, although any linguist can obviously argue for linguistic change in spite of major genetic continuity. But then we are stuck in the pre-ancient DNA era, so what’s ancient DNA for.

    netted-ware-textile-ceramics
    Middle Bronze Age cultures of Eastern Europe.

    Genetic continuity = language continuity?

    In the end, it’s very difficult to say how much language continuity there is around Estonia since the arrival of Corded Ware peoples. Looking at Modern Estonians, they have been clearly influenced by recent contacts with Baltic- and Germanic-speaking peoples clustering to the south-west in the PCA. They seem to have also received contacts from north(-east)ern peoples, likely from Finland, evidenced by their shifts toward the modern Estonian cluster during and after the Middle Ages, with a slight increase in Siberian ancestry and N1c subclades associated with Lovozero Ware. How much language change did these contacts bring? Maybe an expansion of Gulf of Finland Finnic (Northern Estonian) over Inland Finnic (Southern Estonian) and Gulf of Riga Finnic (Livonian)? Difficult to know, exactly, but, in the traditional view of Balto-Finnic dialectal distribution among Uralicists like Kallio, possibly no change at all.

    So, if the obvious changes in the Estonia_MA cluster relative to Estonia_IA cluster and Estonia_Modern relative to Estonia_MA do not represent radical language change…Why would Estonia_IA represent a change relative to Estonia_BA, when it is statistically basically the same? Or Estonia_BA relative to CWC_Baltic? Because of the infiltration of haplogroup N1c around the whole Baltic? Because of the occasional 1% “Siberian” ancestry in some non-locals of varied haplogroups across the whole Baltic area?

    In spite of all this, the amount of special pleading we are seeing among openly Nordicist amateurs when discussing the Uralic homeland relative to the Indo-European question in genetics has become a matter of plain willful ignorance. Like the living corpses of the Anatolian homeland, the Armenian homeland, the OIT proponents, or the nativist Basque R1b association, the personal involvement in the revival of “R1a=Indo-European” and “N=Uralic” trends is just painful to watch.

    [Next post in this line, if I manage to make time for it: “Genetic (dis)continuity in Central Europe“. Let’s see if early Balts and early Slavs, as well as Germanic peoples, show a cluster closer to Danubian EBA (viz. Maros), Hungary-Balkans BA, and Urnfield-related samples than their predecessors in their areas, i.e. away from East Corded Ware groups… If you want, you can enjoy for the moment the new PCAs I could get done and the tentative map of languages in the Early Bronze Age, that will probably give you the right idea about early Indo-European and Uralic population movements]

    bronze-age-early-indo-european
    European Early Bronze Age: tentative language map based on linguistics, archaeology, and genetics. See full map.

    Related

    Baltic Finns in the Bronze Age, of hg. R1a-Z283 and Corded Ware ancestry

    estonian-bronze-age-dna

    Open access The Arrival of Siberian Ancestry Connecting the Eastern Baltic to Uralic Speakers further East, by Saag et al. Current Biology (2019).

    Interesting excerpts:

    In this study, we present new genomic data from Estonian Late Bronze Age stone-cist graves (1200–400 BC) (EstBA) and Pre-Roman Iron Age tarand cemeteries (800/500 BC–50 AD) (EstIA). The cultural background of stone-cist graves indicates strong connections both to the west and the east [20, 21]. The Iron Age (IA) tarands have been proposed to mirror “houses of the dead” found among Uralic peoples of the Volga-Kama region [22].

    (…) The 33 individuals included 15 from EstBA, 6 from EstIA, 5 from Pre-Roman to Roman Iron Age Ingria (500 BC–450 AD) (IngIA), and 7 from Middle Age Estonia (1200–1600 AD) (EstMA) and yielded endogenous DNA ∼4%–88%, average genomic coverages ∼0.017–0.734×, and contamination estimates <4% (Table S1). We analyzed the data in the context of modern and other ancient individuals, including from Neolithic Estonia [13].

    estonian-y-dna-bronze-iron-age
    Archaeological Information, Genetic Sex, mtDNA and Y Chromosome Haplogroups, and Average Coverage of the Individuals of This Study. Modified from the paper to mark distinct Y-DNA haplogroups in the LBA and IA.

    We identified chrY hgs for 30 male individuals (Tables 1 and S2; STAR Methods). All 16 successfully haplogrouped EstBA males belonged to hg R1a, showing no change from the CWC period, when this was also the only chrY lineage detected in the Eastern Baltic [11, 13, 30, 31]. Three EstIA and two IngIA individuals also belonged to hg R1a, but three EstIA males belonged to hg N3a, the earliest so far observed in the Eastern Baltic. Three EstMA individuals belonged to hg N3a, two to hg R1a, and one to hg J2b. ChrY lineages found in the Baltic Sea region before the CWC belong to hgs I, R1b, R1a5, and Q [10, 11, 12, 13, 17, 32]. Thus, it appears that these lineages were substantially replaced in the Eastern Baltic by hg R1a [10, 11, 12, 13], most likely through steppe migrations from the east [30, 31]. (…) Our results enable us to conclude that, although the expansion time for R1a1 and N3a3′5 in Eastern Europe is similar [25], hg N3a likely reached Estonia or at least became comparably frequent to modern Estonia [1] only during the BA-IA transition.

    A clear shift toward West Eurasian hunter-gatherers is visible between European LN and BA (including Baltic CWC) and EstBA individuals, the latter clustering together with Latvian and Lithuanian BA individuals [11]. EstIA, IngIA, and EstMA individuals project between BA individuals and modern Estonians, partially overlapping with both.

    (…) EstBA individuals are clearly distinguishable from Estonian CWC individuals as the former have more of the blue component most frequent in WHGs and less of the brown and yellow components maximized in Caucasus hunter-gatherers and modern Khanty, respectively. The individuals of EstBA, EstIA, IngIA, EstMA, and modern Estonia are quite similar to each other on average, indicating that the relatively high proportion of WHG ancestry in modern Eastern Baltic populations compared to other present-day Europeans [15] traces back to the BA.

    estonian-pca-published
    Detail of the PCA, modified from the paper to label populations. Estonian Bronze Age and Iron Age samples cluster close to Early Corded Ware from the Baltic.. Principal-component analysis results of modern West Eurasians with ancient individuals projected onto the first two components (PC1 and PC2). BA, Bronze Age; EF, early farmers; HG, hunter-gatherers; IA, Iron Age; IMA, Iron/Middle Ages; LN, Late Neolithic; LNBA, Late Neolithic/Bronze Age; MA, Middle Ages

    When comparing Estonian CWC and EstBA using autosomal outgroup f3 and Patterson’s D statistics (Table S3), the latter is more similar to other Baltic BA populations, to Baltic IA and Middle Age (MA) populations, and also to populations similar to WHGs and Scandinavian hunter-gatherers (SHGs), but not to Estonian CCC (Figures 2A and S2A; Data S1). The increase in WHG or SHG ancestry could be connected to western influences seen in material culture [20, 21] and facilitated by a decline in local population after the CCC-CWC period [20]. A slight trend of bigger similarity of Estonian CWC to forest or steppe zone populations and of EstBA to European early farmer populations can also be seen.

    (…) When comparing to modern populations, Estonian CWC is slightly more similar to Caucasus individuals but EstBA to Baltic populations and Finnic speakers (Figure 2B; Data S1). Outgroup f3 and D statistics do not reveal apparent differences when comparing EstBA to EstIA, EstIA to IngIA, and EstIA to EstMA (Data S1).

    estonian-ba-ia-ancestry
    qpAdm results. Error bars indicate one SE. Central MN, Central European Middle Neolithic; EstBA, Estonian Bronze Age; EstIA, Estonian Iron Age; IngIA, Ingrian Iron Age; EstMA, Estonian Middle Ages; WHG, western hunter-gatherers.

    These results highlight how uniparental and autosomal data can lead to different demographic inferences—the genetic change between CWC and BA not seen in uniparental lineages is clear in autosomal data and the appearance of chrY hg N in the IA is not matched by a clear shift in autosomal profiles.

    EstBA individuals have no Nganasan-related ancestry and EstIA, IngIA, and EstMA individuals on average have 2% or 4% (Figure 3; Data S1). The differentiation remains when using BA or IA Fennoscandian populations [26] instead of Nganasans (Data S1). Notably, the proportion of Nganasan-related ancestry varies between 0% and 12% among sampled EstIA, IngIA, and EstMA individuals (Data S1), which may suggest its relatively recent admixture into the target population. Moreover, two individuals from Kunda (0LS10 and V10) have the highest proportions of Nganasan ancestry among EstIA (6% and 8%), one of them has chrY hg N3a, and isotopic analysis suggests neither individual being born in Kunda [34].

    About these two males from Tarand-graves, ‘foreign’ to Kunda:

    0LS10: Male from tarand III (burial 9; TÜ 1325: L777), age 17–25 years [34]. He had a fragment of a sheep/goat bone and ceramics as grave goods. This burial has two radiocarbon dates: 2430 ± 35 BP (Poz-10801; 760–400 cal BC) and 2530 ± 41 BP (UBA-26114; 800–530 cal BC) [34]. According to the isotopic analysis, the person was not born in the vicinity of Kunda; his place of birth is still unknown (but south-western Finland and Sweden are excluded) [34]. Sampled tooth r P1.

    V10: Male from tarand XI (burial 24; TÜ 1325: L1925), age 25–35 years [34], date 2484 ± 40 BP (UBA-26115; 790–430 cal BC) [34]. He had a few potsherds near the skull. Likewise, this person was not locally born [34]. Sampled tooth l P1.

    estonia-bronze-iron-age-steppe-siberian
    Autosomal Analyses’ Results for Gyvakarai1 as the closest available Corded Ware source for Balto-Finnic populations.

    The paper shows thus:

    • Major continuity of ancestry from Corded Ware to modern Estonians, with only slight changes in different periods. In fact, one of the best fits for the Late Bronze Age ancestry is Gyvakarai1, one of the Corded Ware “outliers” described as “closer to Yamna”, which I already said may be closer to Sredni Stog/EHG populations instead. Another interesting take is that the change from Bronze Age to Iron Age corresponds to an increase in Baltic Corded Ware-related ancestry, rather than being driven by Siberian ancestry.
    • pca-mittnik-gyvakarai
      File modified by me from Mittnik et al. (2018) to include the approximate position of the most common ancestral components, and an identification of potential outliers. Zoomed-in version of the European Late Neolithic and Bronze Age samples. “Principal components analysis of 1012 present-day West Eurasians (grey points, modern Baltic populations in dark grey) with 294 projected published ancient and 38 ancient North European samples introduced in this study (marked with a red outline). From Mittnik et al. (2018).
    • A Volosovo-related migration of hg. N1c with Netted Ware into the area seems to be discarded, based on the full replacement of paternal lines and continuity of R1a-Z283. It is only during the Tarand-grave period when a system of chiefdoms (spread from Ananyino/Akozino) brings haplogroup N1c to the Gulf of Finland. During the Iron Age, the proportion of paternal lineages is still clearly in favour of R1a (50% in the coast, 100% in Ostrobothnia), which indicates a gradual replacement led by elites, likely because of the incorporation of Akozino warrior-traders spreading all over the Baltic, bringing the described shared Mordvinic traits in Fennic.
    • finno-ugric-haplogroup-n
      Map of archaeological cultures in north-eastern Europe ca. 8th-3rd centuries BC. [The Mid-Volga Akozino group not depicted] Shaded area represents the Ananino cultural-historical society. Fading purple arrows represent likely stepped movements of subclades of haplogroup N for centuries (e.g. Siberian → Ananino → Akozino → Fennoscandia [N-VL29]; Circum-Arctic → forest-steppe [N1, N2]; etc.). Blue arrows represent eventual expansions of Uralic peoples to the north. Modified image from Vasilyev (2002).
    • The arrival of Akozino warrior-traders (bringing N1c and R1a lineages) was probably linked to this minimal “Nganasan-like” ancestry of some samples in the transition to the Iron Age. This arrival is supported by samples 0LS10 (the earliest hg. N1c) and V10 (of hg. R1a), both dated to ca. 800-400 BC, with V10 showing the highest “Nganasan-like” ancestry with 4.8%, both of them neighbouring samples showing 0%. This variable admixture among local and foreign paternal lineages might support the described social system of family alliances with intermarriages. In fact, a medieval sample, 0LS03_1 (hg. R1a) also shows a recent “Nganasan-like” ancestry, which probably points to the integration of different Arctic-related ancestry components among Modern Estonians, in this case related to Finnish expansions and thus integration of Levänluhta-related ancestry, as per the supplementary data.
    • NOTE. Such minimal proportions of “Nganasan-like” ancestry evidence the process of admixture of Volga Finns in Akozino territory through their close interactions with Permians of Ananyino, who in turn acquired this Palaeo-Arctic admixture most likely during the expansion of the linguistic community to hunter-gatherer territories, to the north of the Cis-Urals. This process of stepped infiltration and expansion without language change is not dissimilar to the one seen among Indo-Iranians and Balto-Slavs of hg. R1b, or Vasconic speakers of hg. I2a, although in the case of Baltic Finns of hg. R1a the process of infiltration and expansion of hg. N1c is much less dramatic, with no radical replacement anywhere before the huge bottlenecks observable in Finns.

    • The expansion of haplogroup N1c among Finnic populations, as we are going to see in samples from the Middle Ages such as Luistari, is the consequence of late founder effects after huge bottlenecks expected based on the analysis of modern populations. The expansion of N1c-VL29 is different in origin from that of N1c-Z1936 among Samic (later integrated into Finnish populations), most likely from the east and originally associated with Lovozero Ware.
    haplogroup_n3a3
    Frequency-Distribution Maps of Individual Subclade N3a3 / N1a1a1a1a1a-CTS2929/VL29, probably initially with Akozino warrior-traders. Map from Ilumäe et al. (2016).

    In spite of all this, the conclusion of the paper is (surprise!) that Siberian ancestry and hg. N heralded the arrival of Finnic to the Gulf of Finland in the Iron Age… However, this conclusion is supposedly* supported, not by their previous papers, but by a recent phylogenetic study by Honkola et al. (2013), which doesn’t actually argue for such a late ‘arrival’: it argues for the split of Balto-Finnic around 1500 BC.

    NOTE. I say ‘supposedly’ because Kristiina Tambets, for example, has been following the link of Uralic with haplogroup N since the 2000s, so this is not some conclusion they just happened to misread from some random paper they Googled. In those initial assessments, she argued that the “ancient homeland” of the Tat C mutation suggested that Finno-Ugrians were in Fennoscandia before Indo-Europeans. Apparently, since haplogroup N appears later and from the east, it is now more important to follow this haplogroup than what is established in archaeology and linguistics.

    Even in the referred paper, this split is considered an in situ development, since the phylogenetic study takes the information – among others – 1) from Parpola and Carpelan, who consider Netted Ware, a culture derived from Fatyanovo/Abashevo and Volosovo, as the culprit of the Finno-Ugric expansion; and 2) from Kallio (2006), who clearly states that Proto-Balto-Finnic (like Proto-Finno-Samic) was spoken around the Gulf of Finland during the Bronze Age. Both of them set the terminus ante quem of the language presence in the Baltic ca. 1900 BC.

    Anyways, as a consequence of geneticists keeping these untenable pre-ancient DNA haplogroup-based arguments today, I expect to see this “Finnic” language expansion also described for the Western Baltic, Scandinavia or northern Europe, when this same proportion of hg. N1c and “Nganasan” ancestry is observed in Iron Age samples around the Baltic Sea. The nativist trends that this domination of “Finns” all over Northern Europe 2,500 years ago will create will be even more fun to read than the current ones…

    EDIT (10 May 2019) How I see the reaction of many to ancient DNA, in keeping their old theories:

    Related

    The cradle of Russians, an obvious Finno-Volgaic genetic hotspot

    pskov-novgorod-russia

    First look of an accepted manuscript (behind paywall), Genome-wide sequence analyses of ethnic populations across Russia, by Zhernakova et al. Genomics (2019).

    Interesting excerpts:

    There remain ongoing discussions about the origins of the ethnic Russian population. The ancestors of ethnic Russians were among the Slavic tribes that separated from the early Indo-European Group, which included ancestors of modern Slavic, Germanic and Baltic speakers, who appeared in the northeastern part of Europe ca. 1,500 years ago. Slavs were found in the central part of Eastern Europe, where they came in direct contact with (and likely assimilation of) the populations speaking Uralic (Volga-Finnish and Baltic- Finnish), and also Baltic languages [11–13]. In the following centuries, Slavs interacted with the Iranian-Persian, Turkic and Scandinavian peoples, all of which in succession may have contributed to the current pattern of genome diversity across the different parts of Russia. At the end of the Middle Ages and in the early modern period, there occurred a division of the East Slavic unity into Russians, Ukrainians and Belarusians. It was the Russians who drove the colonization movement to the East, although other Slavic, Turkic and Finnish peoples took part in this movement, as the eastward migrations brought them to the Ural Mountains and further into Siberia, the Far East, and Alaska. During that interval, the Russians encountered the Finns, Ugrians, and Samoyeds speakers in the Urals, but also the Turkic, Mongolian and Tungus speakers of Siberia. Finally, in the great expanse between the Altai Mountains on the border with Mongolia, and the Bering Strait, they encountered paleo-Asiatic groups that may be genetically closest to the ancestors of the Native Americans. Today’s complex patchwork of human diversity in Russia has continued to be augmented by modern migrations from the Caucasus, and from Central Asia, as modern economic migrations take shape.

    pskov-novgorod-pca-eurasia-yakut
    Sample relatedness based on genotype data. Eurasia: Principal Component plot of 574 modern Russian genomes. Colors reflect geographical regions of collection; shapes reflect the sample source. Red circles show the location of Genome Russia samples.

    In the current study, we annotated whole genome sequences of individuals currently living on the territory of Russia and identifying themselves as ethnic Russian or as members of a named ethnic minority (Fig. 1). We analyzed genetic variation in three modern populations of Russia (ethnic Russians from Pskov and Novgorod regions and ethnic Yakut from the Sakha Republic), and compared them to the recently released genome sequences collected from 52 indigenous Russian populations. The incidence of function-altering mutations was explored by identifying known variants and novel variants and their allele frequencies relative to variation in adjacent European, East Asian and South Asian populations. Genomic variation was further used to estimate genetic distance and relationships, historic gene flow and barriers to gene flow, the extent of population admixture, historic population contractions, and linkage disequilibrium patterns. Lastly, we present demographic models estimating historic founder events within Russia, and a preliminary HapMap of ethnic Russians from the European part of Russia and Yakuts from eastern Siberia.

    pskov-novgorod-pca-finno-permic
    Sample relatedness based on genotype data. Western Russia and neighboring countries: Principal Component plot of 574 modern Russian genomes. Colors reflect geographical regions of collection; shapes reflect the sample source. Red circles show the location of Genome Russia samples.

    The collection of identified SNPs was used to inspect quantitative distinctions among 264 individuals from across Eurasia (Fig. 1) using Principal Component Analysis (PCA) (Fig. 2). The first and the second eigenvectors of the PCA plot are associated with longitude and latitude, respectively, of the sample locations and accurately separate Eurasian populations according to geographic origin. East European samples cluster near Pskov and Novgorod samples, which fall between northern Russians, Finno-Ugric peoples (Karelian, Finns, Veps etc.), and other Northeastern European peoples (Swedes, Central Russians, Estonian, Latvians, Lithuanians, and Ukrainians) (Fig. 2b). Yakut individuals map into the Siberian sample cluster as expected (Fig. 2a). To obtain an extended view of population relationships, we performed a maximum likelihood-based estimation of ancestry and population structure using ADMIXTURE [46](Fig. 2c). The Novgorod and Pskov populations show similar profiles with their Northeastern European ancestors while the Yakut ethnic group showed mixed ancestry similar to the Buryat and Mongolian groups.

    pskov-novgorod-yakut-admixture
    Population structure across samples in 178 populations from five major geographic regions (k=5). Samples are pooled across three different studies that covered the territory of Russian Federation (Mallick et al. 2016 [36], Pagani et al. 2016 [37], this study). The optimal k-value was selected by value of cross validation error. Russian samples from all studies (highlighted in bold dark blue) show a slight gradient from Eastern European (Ukrainian, Belorussian, Polish) to North European (Estonian Karelian, Finnish) structures, reflecting population history of northward expansion. Yakut samples from different studies (highlighted in bold red) also show a slight gradient from Mongolian to Siberian people (Evens), as expected from their original admixture and northward expansions. The samples originated from this study are highlighted, and plotted in separated boxes below.

    Possible admixture sources of the Genome Russia populations were addressed more formally by calculating F3 statistics, which is an allele frequency-based measure, allowing to test if a target population can be modeled as a mixture of two source populations [48]. Results showed that Yakut individuals are best modeled as an admixture of Evens or Evenks with various European populations (Supplemental Table S4). Pskov and Novgorod showed admixture of European with Siberian or Finno-Ugric populations, with Lithuanian and Latvian populations being the dominant European sources for Pskov samples.

    direction-expansion-russians
    The heatmaps of gene flow barriers show for each point at the geographical map the interpolated differences in allele frequencies (AF) between the estimated AF at the point with AFs in the vicinity of this point. The direction of the maximal difference in allele frequencies is coded by colors and arrows.

    So, Russians expanding in the Middle Ages as acculturaded Finno-Volgaic peoples.

    Or maybe the true Germano-Slavonic™-speaking area was in north-eastern Europe, until the recent arrival of Finno-Permians with the totally believable Nganasan-Saami horde, whereas Yamna -> Bell Beaker represented Vasconic-Caucasian expanding all over Europe in the Bronze Age. Because steppe ancestry in Fennoscandia and Modern Basques in Iberia.

    A really hard choice between equally plausible models.

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