When Bell Beakers mixed with Eneolithic Europeans: Pömmelte and the Europe-wide concept of sanctuary

pommelte-enclosure

Recent open access paper The ring sanctuary of Pömmelte, Germany: a monumental, multi-layered metaphor of the late third millennium BC, by Spatzier and Bertemes, Antiquity (2018) 92(363):655-673.

Interesting excerpts (emphasis mine):

In recent decades, evidence has accumulated for comparable enclosures of later dates, including the Early Bronze Age Únětice Culture between 2200 and 1600 BC, and thus into the chronological and cultural context of the Nebra sky disc. Based on the analysis of one of these enclosure sites, recently excavated at Pömmelte on the flood plain of the Elbe River near Magdeburg, Saxony-Anhalt, and dating to the late third millennium BC

The main occupation began at 2321–2211 cal BC, with the stratigraphically earliest features containing exclusively Bell Beaker finds. Bell Beaker ceramics continue after 2204–2154 cal BC (boundary occupation I/II), although they were probably undecorated, but are now complemented by Únětice Culture (and other Early Bronze Age) types. At this time, with features common to both cultures predominate. Only contexts dating to the late main occupation phase (late phase II) and thereafter contained exclusively Únětice Culture finds. Evidently, the bearers of the Bell Beaker Culture were the original builders of the enclosure. During a second phase of use, Final Neolithic and Early Bronze Age cultures coexisted and intermingled. The material remains, however, should not be taken as evidence for successive groups of differing archaeological cultures, but as witnesses to a cultural transition from the Bell Beaker Culture to the Únětice Culture (Spatzier 2015). The main occupation ended 2086–2021 cal BC with the deconstruction of the enclosure; Bell Beaker finds are now absent. Finally, a few features (among them one shaft) and radiocarbon dates attest the sporadic re-use of the site in a phase of abandonment/re-use that ended 1636– 1488 cal BC.

pommelte-enclosure-occupation-stratigraphy
Cultural sequence and chronological model of the Pömmelte enclosure’s occupation (dates in 1σ-precision) (designed by André Spatzier).

How the above-ground structures possibly influenced perception may reveal another layer of meaning that highlights social functions related to ritual. While zone I was disconnected from the surroundings by a ‘semi-translucent’ post-built border, zones II/III were separated from the outside world by a wooden wall (i.e. the palisade), and zone III probably separated individuals from the crowd gathered in zone II. Accessing the interior or centre therefore meant passing through transitional zones, to first be secluded and then segregated. Exiting the structure meant re-integration and re-connection. The experience possibly induced when entering and leaving the monument reflects the three stages of ‘rites of passage’ described by van Gennep (1909): separation, liminality and incorporation. The enclosure’s outer zone(s) represents the pre- and post-liminal phase; the central area, the liminal phase. Seclusion and liminality in the interior promoted a sense of togetherness, which can be linked to Turner’s “communitas” (1969: 132–33). We might therefore see monuments such as the Pömmelte enclosure as important communal structures for social regulation and the formation of identity.

ring-sanctuary-of-pommelte
Layers of meaning of the Pömmelte enclosure as deduced from the archaeological record (design by André Spatzier).

(…) The long-term stability of these connotations must be emphasised. As with the tradition of making depositions, these meanings were valid from the start of the occupation — c. 2300 BC — until at least the early period following the deconstruction event, c. 2050 BC. While the spatial organisation and the solar alignment of the main entrances were maintained throughout the main occupation, stone axes and ‘formal’ graves indicate the continuation of the spatial concepts described above until the twentieth to nineteenth centuries BC.

These layers of meaning mirror parallel concepts of space including, although not necessarily restricted to, the formation of group identities (see Hansen & Meyer 2013: 5). They can perhaps be better understood as a ‘cosmological geography’ manifested in the symbolism of superimposed levels of conceptual ideas related to space and to certain cardinal points (Figure 8). This idea is closely related to Eliade’s (1959: 29–36) understanding of “organized — hence comicized — territory”, that is territory consecrated to provide orientation within the homogeneity of the chaotic ‘outside world’, and the equivalence of spatial consecration and cosmogony. Put differently, the Pömmelte enclosure can be interpreted as a man-made metaphor and an icon of the cosmos, reflecting the Weltanschauung (a comprehensive conception of the world) of the people who built and used it. By bringing together Eliade and Rappaport’s ideas of meaningfulness in relation to religious experience (Rappaport 1999: 391–95), it may be argued that Pömmelte was a place intended to induce oneness with the cosmos. In combining multiple layers that symbolically represent different aspects of life (first-ordermeaning), the enclosure became an icon metaphorically representing the world (second-order-meaning). As this icon was the place to reaffirm life symbolism ritually, through their actions, people perhaps experienced a sense of rootedness in, or unity with, the cosmos (highest-order-meaning). Although we can only speculate about the perceptions of ancient people, such a theory aiming to describe general principles of religious experience can provide insight.

Conclusions

The circular enclosure of Pömmelte is the first Central European monumental complex of primarily sacred importance that has been excavated and studied in detail. It reveals aspects of society and belief during the transition from the Final Neolithic to the Early Bronze Age, in the second half of the third millennium BC. Furthermore, it offers details of ritual behaviour and the way that people organised their landscape. A sacred interior was separated from the profane environment, and served as a venue for rites that secured the continuity of the social, spiritual and cosmic order. Ancestor worship formed another integral part of this: a mound-covered burial hut and a square-shaped ditch sanctuary (located, respectively, within and near the enclosure’s south-eastern sector; cf. Figure 2)—dating to 2880–2580 cal BC and attributed to the Corded Ware Culture (Spatzier 2017a: 235–44)—suggest that this site was deliberately chosen. With construction of the ring sanctuary, this place gained an immense expansion in meaning—comparable to Stonehenge. Through architectural transformation, both of these sites developed into sanctuaries with increasingly complex religious functions, including in relation to the cult of the dead. The cosmological and social functions, and the powerful symbolism of the Nebra sky disc and hoard (Meller 2010: 59–70), are reflected in Pömmelte’s monumental architecture.

All of these features—along with Pömmelte’s dating, function and complex ring structure—are well documented for British henge monuments (Harding 2003; Gibson 2005). The continuous use of circular enclosures in Central Europe from around 3000– 1500 BC remains to be confirmed, but strong evidence indicates usage spanning from the fifth to the first millennia BC (Spatzier 2017a: 273–96). From 2500 BC onwards, examples in Central Europe, Iberia and Bulgaria (Bertemes 2002; Escudero Carrillo et al. 2017) suggest a Europe-wide concept of sanctuary. This indicates that in extensive communication networks at the beginning of bronze metallurgy (Bertemes 2016), intellectual and religious contents circulated alongside raw materials. The henge monuments of the British Isles are generally considered to represent a uniquely British phenomenon, unrelated to Continental Europe; this position should now be reconsidered. The uniqueness of Stonehenge lies, strictly speaking, with its monumental megalithic architecture.

pommelte-enclosure-space
Model of the spatial organisation of the Pömmelte enclosure (designed by André Spatzier).

The Classical Bell Beaker heritage

No serious scholar can argue at this point against the male-biased East Bell Beaker migrations that expanded the European languages related to Late Proto-Indo-European-speaking Yamna (see David Reich’s comments), and thus most likely North-West Indo-European – the ancestor of Italo-Celtic, Germanic, and Balto-Slavic, apart from Pre-Celtic IE in the British Isles, Lusitano-Galician in Iberia, or Messapic in Italy (see here a full account).

With language, these migrants (several ten thousands) brought their particular Weltanschauung to all of Western, Central, and Northern Europe. Their admixture precisely in Hungary shows that they had close interactions with non-Indo-European peoples (genetically related to the Globular Amphorae culture), something that we knew from the dozens of non-Indo-European words reconstructed exclusively for North-West Indo-European, apart from the few reconstructed non-Indo-European words that NWIE shares with Palaeo-Balkan languages, which point to earlier loans from their ancestors, Yamna settlers migrating along the lower Danube.

It is not difficult to imagine that the initial East Bell Beaker group shared a newly developed common cosmological point of view that clashed with other neighbouring Yamna-related worldviews (e.g. in Balkan EBA cultures) after the cultural ties with Yamna were broken. Interesting in this respect is for example their developed (in mythology as in the new North-West Indo-European concept) *Perkwūnos, the weather god – probably remade (in language as in concept) from a Yamna minor god also behind Old Indian parjányas, the rain god – as one of the main gods from the new Pantheon, distinct from *Dyēus patēr, the almighty father sky god. In support of this, the word *meldh-n- ‘lightning’, behind the name of the mythological hammer of the weather god (cf. Old Norse Mjǫllnir or Latvian Milna), was also a newly coined North-West Indo-European term, although the myth of the hero slaying the dragon with the magical object is older.

perkunos-perkunas
The Hand of Perkūnas by Mikalojus Konstantinas Čiurlionis, from Wikipedia

Circular enclosures are known in Europe since the Neolithic. Also, the site selected for the Pömmelte enclosure had been used to bury Corded Ware individuals some centuries before its construction, and Corded Ware symbolism (stone axe vs. quern) is seen in the use given by Bell Beakers and later Únětice at this place. All this and other regional similarities between Bell Beakers and different local cultures (see here an example of Iberian Bell Beakers) points to syncretism of the different Bell Beaker groups with preceding cultures in the occupied regions. After all, their genealogical ancestors included also those of their maternal side, and not all encountered males disappeared, as is clearly seen in the resurge of previous paternal lineages in Central-East Europe and in Scandinavia. The admixture of Bell Beakers with previous groups (especially those of similar steppe-related ancestry from Corded Ware) needs more complex analyses to clarify potential early dialectal expansions (read what Iosif Lazaridis has to say).

The popular “big and early” expansions

These syncretic trends gave rise to distinct regional cultures, and eventually different local groups rose to power in the new cultural regions and ousted the old structures. Social norms, hierarchy, and pantheons were remade. Events like this must have been repeated again and again in Bronze and Iron Age Europe, and in many cases it was marked by a difference in the prevailing archaeological culture attested, and probably accompanied by certain population replacements that will be seen with more samples and studies of fine-scale population structure.

Some of these cultural changes, marked by evident haplogroup or admixture replacement, are defined as a ‘resurge’ of ancestry linked to previous populations, although that is obviously not equivalent to a resurge of a previous cultural group, because they usually represent just a successful local group of the same supraregional culture with a distinct admixture and/or haplogroup (see e.g. resurge of R1a-Z645 in Central-East European Bronze Age). Social, religious, or ethnic concepts may have changed in each of these episodes, along with the new prestige dialect.

NOTE. A recent open access paper on two newly studied Middle Bronze Age inhumations from Stonehenge give an interesting idea of potential differences in social identities, in ancestry and geographic origin (which characterize ethnicity) may have been marked by differences in burial ceremonies: Lives before and after Stonehenge: An osteobiographical study of four prehistoric burials recently excavated from the Stonehenge World Heritage Site, by Mays et al. Journal of Archaeological Science: Reports (2018) 20:692-710.

This must have happened then many times during the hundreds (or thousands in some cases) of years until the first attestation of a precise ancient language and culture (read e.g. about one of the latest branches to be attested, Balto-Slavic). Ancient language contacts, like substrates or toponymy, can only rarely be detected after so many changes, so their absence (or the lack of proper studies on them) is usually not relevant – and certainly not an argument – in scholarly discussions. Their presence, on the other hand, is a proof of such contacts.

chalcolithic_late_Europe_Bell_Beaker
Diachronic map of Late Copper Age migrations including Classical Bell Beaker (east group) expansion from central Europe ca. 2600-2250 BC

We have dozens of papers supporting Uralic dialectal substrate influence on Pre-Germanic, Proto-Balto-Slavic, and Pre- and Proto-Indo-Iranian (and even Proto-Celtic), as well as superstrate influence of Palaeo-Germanic (i.e. from Pre- to Proto-Germanic) and Proto-Balto-Slavic into Proto-Finno-Saamic, much stronger than the Indo-Iranian adstrate influence on Finno-Ugric (see the relative importance of each influence) which locates all these languages and their evolution to the north and west of the steppe (with Proto-Permic already separated, in North-East Europe, as is Proto-Ugric further east near the Urals), probably around the Baltic and Scandinavia after the expansion of Bell Beakers. These connections have been known in linguistics for decades.

Apart from some early 20th century scholars, only a minority of Indo-Europeanists support nowadays an Indo-European (i.e. centum) substrate for Balto-Slavic, to keep alive an Indo-Slavonic group based on a hypothetical 19th century Satem group; so e.g. Holzer with his Temematic, and Kortlandt supporting him, also with some supposed Indo-European substrate with heavy non-Indo-European influence for Germanic and Balto-Slavic, that now (thanks mainly to the views of the Copenhagen group) have been linked to the Corded Ware culture, as it has become clear even to them that Bell Beakers expanded North-West Indo-European.

NOTE. The Temematic etymologies have been (all of them) fully dismissed e.g. in Matasović (2013). I have already explained why an Indo-Slavonic group from Sredni Stog is not tenable, and genetics (showing Late PIE only from Yamna expansions) is proving that, too.

For their part, only a minority among Uralicists, such as Kuz’mina, Parpola or Häkkinen, believe in an ‘eastern’ origin of Uralic languages, around the Southern Urals. Genomic finds – like their peers – are clearly not supporting their views. But even if we accept this hypothesis, there is little space beyond Abashevo and related East Corded Ware cultures after the recent papers on Corded Ware and Fennoscandian samples. And yet here we are:

The Copenhagen “Homeland” interactive map

copenhagen-group-map
Brought to you by the Copenhagen fantasy map series, Indo-Europeans after (no, really, after) the expansion of Yamna settlers in Hungary ca. 2700 BC: Yamna settlers have magically disappeared. Yamna-related Balkan EBA cultures and the hundreds of Yamna kurgans around the Lower Danube and in Hungary up to Saxony-Anhalt do not exist. Dat huge mythical Middle Dnieper territory lasting (unchanged) for a thousand years, in sooo close contact with Yamna territory (so beautifully ‘linked’ together that they must have been BFFs and admixed!). Uralic Mesolithic hunter-gatherers resisting IE invasions in Volosovo for 1,500 years like Asterix’ Gaulish village against the Romans. Tiny pockets of Bell Beakers will eventually emerge from (surprise!) Corded Ware territories beautifully scattered over Central and Northern Europe (unlike those eastern CWC mega-regions). And, of course, you can almost see Kroonen & Iversen’s Kurgan Pre-Germanic mixing already with their agricultural substrate TRB precisely in full-IE Denmark (quite appropriate for the Danish school). And sheep symbols representing wool finds, for no reason. A great map to mock for years to come, with each new genetic paper.

The new propaganda tool GIS timeline map of the Copenhagen group:

  • consciously ignores Yamna settlers along the Danube, in the Balkans, and in Hungary, and initial East Bell Beakers, i.e. the obvious origin and expansion of North-West Indo-Europeans, but in contrast magnifies (and expands in time) regions for Sredni Stog / Corded Ware cultures (which suggests that this is yet another absurd attempt to revive the theories of the Danish school…);
  • substitutes arrows for Kron-like colors (where danger red = Indo-European) with the same end result of many other late 20th century whole-Europe Kurgan maps, linking Sredni Stog and Corded Ware with Yamna, but obviating the precise origin of Corded Ware peoples (is it Sredni Stog, or is it that immutable Middle Dnieper group? is it West Yamna, or Yamna Hungary? is it wool, or is it wheels?);
  • relegates Uralic speakers to a tiny corner, a ‘Volosovo’ cultural region, thus near Khvalynsk/Yamna (but not too much), that miraculously survives surrounded by all-early-splitting, all-Northern Eneolithic Indo-Europeans, thus considering Uralic languages irrelevant not only to locate the PIE Urheimat, but also to locate their own homeland; also, cultures identified in color with Uralic speakers expand until the Iron Age with enough care not to even touch in the map one of the known R1a samples published to date (because, for some people, apparently R1a must be Indo-European); and of course N1c or Siberian ancestry are irrelevant, too;
  • and adds findings of wheels and wool probably in support of some new ideas based on yet another correlation = causation argument (that I cannot then properly criticize without access to its reasoning beyond cute SmartArt-like symbols) similar to their model – already becoming a classic example of wrong use of statistical methods – based on the infamously named Yamnaya ancestral component, which is obviously still used here, too.

The end result is thus similar to any other simplistic 1990s Gimbutas (or rather the recently radicalized IE Sredni Stog -> Corded Ware -> BBC version by the Danish workgroup) + 2000s R1a-map + 2010s Yamnaya ancestry; but, hard to believe, it is published in mid-2018. A lot of hours of senseless effort, because after its publication it becomes ipso facto outdated.

For comparison of Yamna and Bell Beaker expansions, here is a recent simplistic, static (and yet more accurate) pair of maps, from the Reich Lab:

corded-ware-bell-beaker
Cultural maps from Eneolithic and Chalcolithic cultures in Wang et al. (2018).

If the Copenhagen group keeps on pushing Gimbutas’ long ago outdated IE Sredni Stog -> Corded Ware theory as modified by Kristiansen, with their recently invented Corded Ware -> Bell Beaker model in genetics, at some point they are bound to clash with the Reich-Jena team, which seems to have less attachment to the classic Kurgan model and the wrong interpretations of the 2015 papers, and that would be something to behold. Because, as Cersei would say: “When you play the game of thrones, you win or you die. There is no middle ground.” And when you play the game of credibility, after so many, so wrong publications, well…

NOTE. I have been working on a similar GIS tool for quite some time, using my own maps and compiled genetic data, which I currently only use for my 2018 revision of the Indo-European demic diffusion model. Maybe within some weeks or months I will be able to publish the maps properly, after the revised papers. It’s a pitty that so much work on GIS and analysis with genetic data and cultural regions has to be duplicated, but I intend to keep some decent neutrality in my revised cultural maps, and this seems impossible at this point with some workgroups who have put all their eggs in one broken basket…

Related

About Scepters, Horses, and War: on Khvalynsk migrants in the Caucasus and the Danube

steppe-horse-sceptre-khvalynsk

dergachev-scepters-khavlynsk-horsesAbout two months ago I stumbled upon a gem in archaeological studies related to Proto-Indo-Europeans, the book О скипетрах, о лошадях, о войне: этюды в защиту миграционной концепции М.Гимбутас (On sceptres, on horses, on war: Studies in defence of M. Gimbutas’ migration concepts), 2007, by V. A. Dergachev, from the Institute of Cultural Heritage of the Moldavian Republic.

Dergachev’s work dedicates 488 pages to a very specific Final Neolithic-Eneolithic period in the Pontic-Caspian steppe, and the most relevant parts of the book concern the nature and expansion of horses and horse domestication, horse-head scepters, and other horse-related symbology – arguably the most relevant cultural signs associated with Proto-Indo-European speakers in this period.

I haven’t had enough time to read the whole book, but I have read with interest certain important chapters.

About Scepters

Typological classification

The genetic and chronological relationship of horse-head pommel-scepters is classified with incredible detail, to the extent that one could divide subregions among those cultures using them.

khvalynsk-horse-head-scepters
Scheme of regional distribution – chronological – typological development of the carved horse-head stone scepters.

Simplified conclusions of this section include (emphasis mine):

  1. The [horse-head pommel-]scepters arose originally in the depth of the Khvalynsk culture. Following the now well-known finds, they are definitely related to those of the Middle Volga group.
  2. horse-head-pommel-scepters-distribution
    General scheme of genetic and chronological development of carved scepters by visual assessment of morphological details.
  3. In their next modifications, these scepters continued to evolve and develop into the area of the Khvalynsk culture in its latest stages, and possibly later.
  4. Simultaneously, with the same modifications, these scepters “are introduced” into common usage in the Novodanilovka culture, which in its spread by one wing was in contact and interspersed immediately with the area of Khvalynsk remains; and on the other hand, far in the south – in the Pre-Kuban and Ciscaucasian regions – within the range of the Domaikopska culture; and in the west – in the Carpathian – Post-Kuban – with the areas of early agricultural cultures Cucuteni A – Trypillia B1, Gumelnița-Karanovo VI.
  5. The simultaneous presence in the areas of the Ciscaucasian, Carpatho-Danubian, and especially Novodinilovka cultures, whose carriers continue the Khvalynian traditions of making stone scepters, and the scepters themselves (in their non-functional implication in the local cultural environment), all definitely allow us to view these findings as imported Novodanilovka objects.
distribution-horse-scepters
Schematic depiction of the spread of horse-head scepters in the Middle Eneolithic. See a full version with notes here.

Cultural relevance of scepters

The text goes on to make an international comparison of scepters and their relevance as a cultural phenomenon, with its strong symbolic functions as divine object, its use in times of peace, in times of war, and in a system of ritual power.

horse-scepters-steppe
Restoration of V. A. Dergachev: a) model for restoration – Paleolithic and Neolithic wands; b) the expected appearance of the Eneolithic scepter on the handle with a coupling (according to Dergachev 2007).
Especially interesting is the section dedicated to Agamemnon’s scepter in the Iliad, one of the oldest Indo-European epics. Here is an excerpt from Illiad II.100-110 (see here the Greek version) with the scepter’s human and divine genealogy:

Then among them lord Agamemnon uprose, bearing in his hands the sceptre which Hephaestus had wrought with toil. Hephaestus gave it to king Zeus, son of Cronos, and Zeus gave it to the messenger Argeïphontes; and Hermes, the lord, gave it to Pelops, driver of horses, and Pelops in turn gave it to Atreus, shepherd of the host; and Atreus at his death left it to Thyestes, rich in flocks, and Thyestes again left it to Agamemnon to bear, that so he might be lord of many isles and of all Argos.

About the horse

His studies on horse remains show an interesting, detailed quantitative and statistical approach to the importance and (cultural and chronological) origin of horses (and likely horse domestication) in each culture.

Although the part on horse remains is probably a bit outdated today, after many recent studies of Eneolithic steppe sites (see here one example), it still shows the relative distribution of horse bone remains among different steppe cultures, which is probably similar to what could be reported today:

distribution-horses-steppe-eneolithic
Territorial distribution of horse remains in the Middle Eneolithic period. Absolute and relative numbers.

Even more interesting is the relationship of the distribution of horse remains with archaeological complexes and horse-related symbols. Some excerpts from the conclusions of this section:

  1. Accounting and analysis of archeo-zoological and archaeological data proper for a horse for a vast area from the Tisza and the Middle Danube to the Caucasus and the Urals (which includes the main cultures of the western agricultural, Caucasian, and Eastern European cultural zones) clearly points to the eastern cultural zone as a zone of the originally the most important social significance of a horse as the only possible zone of the earliest domestication, horseback riding and all-round use of a horse. In relation to the eastern, the western land – the ancient Carpatho-Danubian or the Caucasian cultural zones – are secondary and subordinate to the first on the phenomenon under consideration.
  2. horse-symbols
    Horse-shaped hanger-amulets made of bone.
  3. The first quantitative leap in the manifestation of the remnants of a horse, marking itself and the first qualitative changes in the social status of this animal, is due mainly to the Middle Volga culture of the developed Neolithic of the Middle Volga region (in part, the Southwest Urals), which, accordingly, determine the cultural context, time and geographic region – or, the initial, single and main epicenter of the process of taming and domestication of a horse.
  4. On the one hand, the subsequent substantial increase in the number of horse remnants, and, on the other, the wide inclusion of the horse in cults, rituals, funerary rituals (horse pendants, ornamented metacarpus, horse bones, sacrificial altars) in the Samara culture of the Early Eneolithic of the same region definitely indicates the continuing increase in the social significance of this species of animal, which was most likely expressed in the final design of a specialized horse breeding culture and, accordingly, in a wide range of applications using a horse for riding. At the same time, we can observe the beginning of the transfer of the already domesticated horse from the original historical and geographic epicenter to other cultures of the eastern cultural zone and, in part, the cultures closest to the periphery of this zone, into the western agricultural zone (Bolgrad-Aldeni P, Pre-CuCuteni-Trypillya A) .
  5. expansion-horse-steppe
    Schematic depiction of cultures and regional-chronological distribution of percentage of horse remains. (Depicted are arrows from Middle Volga and Samara culture to the rest)
  6. Middle Eneolithic – early stages. One of the leading places in the remnants of the horse is in the Middle Volga region, the Khvalynsk culture. Genetically related to the Samara, the Khvalynsk I culture preserves the traditions of the ritual, cultural meaning, the treatment of the image of a horse in funerals (altars, horse bones, funerary rituals). But, At the same time, it is in this precise culture that the image of the horse, included in the social symbolism (horse-head pommel-scepter), for the first time it acquires a special, maximum social significance. That is why the appearance and subsequent widespread distribution of the social symbols in Novodanilovka-type objects can definitely be considered as another qualitative leap in the social significance of a horse – its use for military purposes for close and distant expeditions. And such an interpretation is fully confirmed from the analysis of Novodanilovka-type objects, which is the subject of discussion.
  7. Judging by the osteological data and the typological evolution of the horse-head scepters, the Khvalynian culture and remains of the Novodanilovka type are already associated with the relatively widespread and intensive findings of domesticated horses in various areas of the eastern cultural zone (semi-desert regions of the Lower Volga and the Caspian region – Khvalynsk culture, forest-steppe and steppe from the Volga to the Dnieper – Sredni Stog, Repin cultures), and the western – agricultural (Gumelnitsa, Cucuteni A-Tripolye Bl), and the Caucasus (Pre-Maykop) zones, where, however, the horse played a very modest role.
  8. samara-khvalynsk-horses
    Schematic depiction of cultures and regional-chronological distribution of zooarchaeological and ritual data on horses. (Shadowed are from top to bottom the Middle Volga, Samara, Khvalynsk, and Novodanilovka; in bold, other percentages of unrelated cultures: e.g. to the left of Khvalynsk and Novodanilovka, Sredni Stog with 29.65% overall horse bone remains, but 0% of horse symbolism)
  9. From the functional point of view, according to the sum of the data, there is no reason to doubt that in the eastern zone the horse is already present in the Late Neolithic period. Since its domestication and the emergence of a specialized horse breeding, it has been also widely used for meat, milk and dairy products (including the traditional hippace tradition of the later Scythians), and since the beginning of the early Eneolithic for transport and for riding purposes. Another thing is the horse as a means of war, a means of distant travel and expansion. The beginning of the use of a horse for these purposes, in the opinion of the author, is determined by the appearance of social symbolism in the form of horse-head scepters, and is most fully reflected in the memories of the Khvalynsk culture and, in particular, the Novodanilovka type. Concerning western or Caucasian cultural zones related to Khvalynsk, the horse is thought to have been linked to the eastern region, used mainly for riding, as a means of transport and for communication, which, however, does not exclude its use for meat.

These are the main conclusions-interpretations, suggesting the analysis and archaeological and other sources containing information about the horse. And as for our pommel-scepters, then, as can be seen from these sources, the main thing is that the culture of the Middle Volga region, according to all the data, definitely accumulates in itself the longest traditions associated with the gradual increase of social significance of the horse. And if so, this circumstance motivates the possibility or necessity of appearing in the environment of the bearers of this culture of unique signs-symbols that carry within themselves or reflect the image of this animal as an extremely significant social reality. The revealed and characterized quality, as a matter of fact, fill or open by themselves the hypothetical elements we have previously identified, the meanings of that particularity, folded in the social sign-symbol, in our case – the horse-head-shaped scepter.

horse-symbolism-rituals-steppe
Archaeological sites with objects (signs-symbols) related to horses. Horse-head scepters included in other maps are excluded from this one (notice the conspicuous absence of such objects in Sredni Stog and neighbouring North Pontic regions).

The relevance of Dergachev’s work

As you certainly know by now if you are a usual reader of this blog, there were two other seminal publications that same year correcting and expanding Gimbutas’ model:

Each one of these works taken independently (especially the books) may give a different version of Proto-Indo-European migrations; Anthony and Dergachev are heirs of Gimbutas’ simplistic kurgan-based model, and of other previous, now rejected ideas, and they reflect them whenever they don’t deal with first-hand investigation (and even sometimes when interpreting their own data). Taken together – and especially in combination with recent genetic studies – , though, they describe a clearer, solider model of how Proto-Indo-Europeans developed and expanded.

distribution-scepters-steppe
Distribution of horse-head scepters, according to Dergachev, Sorokin (1986).

Anthony’s publication overshadowed the importance of Dergachev’s work for the English-speaking world – and by extension for the rest of us. However, V. A. Dergachev’s updated study of his previous work on steppe cultures shows the right, thorough, and diligent way of describing the expansion of early Khvalynsk-Novodanilovka chieftains with the horse and horse symbolism into the Caucasus and the Lower Danube (like the seminal work of Harrison & Heyd 2007 described the expansion of Yamna settlers with East Bell Beakers, culturally opposed to Corded Ware and to the Proto-Beakers). On the other hand, Anthony’s broad-brush, superficial description of thousands of years of potential Indo-European-speaking peoples gave a migration picture that – although generally right (like radiocarbon-based Iberian origin of the Bell Beaker culture was right) – was bound to be wrong in some essential details, as we are seeing in archaeology and genetics.

NOTE. As I have said before, Anthony’s interpretations of Sredni Stog culture representing a sort of ‘peasants’ under the rule of Novodanilovka chiefs was based on old theories of Telegin, who changed his mind – as did the rest of the Russian school well before the publication of Dergachev’s book, considering both as distinct cultural phenomena. Anthony selected the old interpretation, not to follow a Gimbutas / Kristiansen model of Sredni Stog being Indo-European and expanding with GAC into Corded Ware (because, for him, Corded Ware peoples were originally non-Indo-European speakers): he seems to have done it to prove that Proto-Anatolian traveled indeed through the North Pontic area, i.e. to avoid the regional ‘gap’ in the maps, if you like. Then with the expansion of Repin over the area, Sredni Stog peoples would have been absorbed. With genetic investigation, as we know, and with this kind of detailed archaeological studies, the traditional preference for “large and early” IE territories – proper of the mid-20th century – are no longer necessary.

sredni-stog-suvorovo-novodanilovka-cernavoda
Anthony (2007): “Steppe and Danubian sites at the time of the Suvorovo-Novodanilovka intrusion, about 4200-3900 BC.”

Steppe Eneolithic

We already had in 2016 a Samara hunter-gatherer sample dated ca. 5600 BC, representative of EHG ancestry, of haplogroup R1b1a. We also had three early Khvalynsk samples from Samara Eneolithic dated ca. 4600 BC, with a drift towards (what we believe now is) a population from the Caucasus, showing haplogroups Q1a, R1a1(xM198), and R1b1a, the last one described in its paper as from a high-status burial, similar to high-status individuals buried under kurgans in later Yamna graves (of R1b-L23 lineages), and therefore likely a founder of an elite group of patrilineally-related families, while the R1a1 sample showed scarce decoration, and does not belong to the M417 lineage expanded later in Sredni Stog or Corded Ware.

In 2017 we knew of the Ukraine_Eneolithic sample I6561, from Alexandria, of a precise subclade (L657) of haplogroup R1a-Z93, dated ca. 4000 BC, and likely from the Sredni Stog (or maybe Kvitjana) culture. This sample alone makes it quite likely that the expansion of R1a-Z645 subclades happened earlier than expected, and that it was associated with movements along forest-steppe cultures, most likely along the Upper Dniester or Dnieper-Dniester corridor up to the Forest Zone.

We have now confirmation that Khvalynsk samples from the Yekaterinovka Cape settlement ca. 4250-4000 BC were reported by a genetic lab (to the archaeological team responsible) as being of R1b-L23 subclades, although the precise clades (reported as P312 and U106) are possibly not accurate.

NOTE. Curiously enough, and quite revealing for the close relationship of scepters to the ritual source of power for Khvalynsk chieftains (political and/or religious leaders), the scepter found in the elite burial 45 of the Ekaterinovka cape (a riverine settlement) shows a unique zoomorphic carving, possibly resembling a toothed fish or reptile, rather than the most common horse-related motifs of the time.

ekaterinovka-cape-scepter
Zoomorphic carved stone scepter of the Ekaterinovka Cape burial 45: photos (left) and schematic depiction (right).

With Wang et al. (2018), a real game-changer in the Khvalynsk – Sredni Stog (and also in the Yamna/Bell Beaker – Corded Ware) opposition, we also know that two Steppe Eneolithic samples from the Northern Caucasus Piedmont, dated ca. 4300-4100 BC, show haplogroup R1b1. Although its direct connection to the expansion of early Khvalynsk with horse-related symbolism is not clear from the archaeological information shared (none), this is what the paper has to say about them:

The two distinct clusters are already visible in the oldest individuals of our temporal transect, dated to the Eneolithic period (~6300-6100 yBP/4300-4100 calBCE). Three individuals from the sites of Progress 2 and Vonjuchka 1 in the North Caucasus piedmont steppe (‘Eneolithic steppe’), which harbor Eastern and Caucasian hunter-gatherer related ancestry (EHG and CHG, respectively), are genetically very similar to Eneolithic individuals from Khalynsk II and the Samara region19, 27. This extends the cline of dilution of EHG ancestry via CHG/Iranian-like ancestry to sites immediately north of the Caucasus foothills.

In contrast, the oldest individuals from the northern mountain flank itself, which are three first degree-related individuals from the Unakozovskaya cave associated with the Darkveti-Meshoko Eneolithic culture (analysis label ‘Eneolithic Caucasus’) show mixed ancestry mostly derived from sources related to the Anatolian Neolithic (orange) and CHG/Iran Neolithic (green) in the ADMIXTURE plot (Fig. 2C). While similar ancestry profiles have been reported for Anatolian and Armenian Chalcolithic and Bronze Age individuals20, 23, this result suggests the presence of the mixed Anatolian/Iranian/CHG related ancestry north of the Great Caucasus Range as early as ~6500 years ago.

On the specific burials, we have e.g. the recent open access paper New cases of trepanations from the 5th to 3rd millennia BC in Southern Russia in the context of previous research: Possible evidence for a ritually motivated tradition of cranial surgery?, by Gresky et al. J Am Phys Anthropol (2016):

During the late 5th millennium BC, cultural groups of the Eneolithic occupied the northern circumpontic area and the areas between the North Caucasus and the Lower Volga. For the first time, individual inhumations were placed below low burial mounds (Rassamakin, 2011). During the 4th millennium BC, the area split into two cultural spheres. In the northern steppe area communities continued with the burial practice of crouched inhumations below low mounds, with this culturally transforming into the early Pit Grave culture. In contrast, in the Caucasian foothill zone and the neighbouring steppe, the Majkop-Novosvobodnaya culture emerged (Kohl and Trifonov, 2014). Similarly, during the 3rd millennium BC, two cultural spheres influenced the area: The North Caucasian Culture dominated the Caucasian foothills for the next five centuries, while in the steppe area between the Lower Don and the Caucasus, regional groups of the Catacomb Culture existed side-by-side.

Burials of the Eneolithic epoch (late 5th millennium BC)

The oldest group of individuals with trepanations are found in the North Caucasian variant of the late circumpontic Eneolithic and date to the last third of the 5th millennium BC (Korenevsky, 2012). Burials of this epoch are inhumations in shallow pits, chiefly without burial goods, but covered with large quantities of red ochre. Of special interest is a collective burial of seven individuals from VP 1/12, who were interred together in a secondary burial ritual. The sites of Tuzluki, Mukhin, Voinuchka, Progress, and Sengileevskii all belong to this period.

PCA-caucasus-khvalynsk-sredni-stog
Image modified from Wang et al. (2018). Samples projected in PCA of 84 modern-day West Eurasian populations (open symbols). Previously known clusters have been marked and referenced. An EHG and a Caucasus ‘clouds’ have been drawn, leaving Pontic-Caspian steppe and derived groups between them.See the original file here.

Without the datasets to test different models, you can only imagine what is happening with the processed, secondary data we have. The position of Eneolithic Steppe cluster in the PCA (probably Khvalynsk-related peoples already influenced by the absorbed, previous Caucasus population), as well as other potential Caucasus groups intermediate between Steppe Maykop and Caucasus Maykop (as suggested by other ancient and modern Caucasus samples), may indicate that Yamna is between Khvalynsk and such intermediate Caucasus populations (as the source of the additional CHG-related ancestry) and – as the paper itself states – that it also received additional EEF contribution, probably from the western cultures absorbed during these Khvalynsk-Novodanilovka migrations (or later during Khvalynsk/Repin migrations).

Also interpreted in light of these early Khvalynsk-Novodanilovka migrations of horse riding chieftains (and their close contacts with the Caucasus), you can clearly see where the similar CHG-like contribution to Ukraine Eneolithic and other North Pontic forest-steppe cultures (which later contributed to Proto-Corded Ware peoples) must have come from. The simplistically reported proportions of EHG:CHG:EEF ancestry might be similar in many of these groups, but the precise origin and evolution of such ancestral components is certainly not the same: statistical methods will eventually show this, when (and if) we have many more samples, but for the moment Y-DNA is the most obvious indicator of such differences.

There was no steppe people speaking a steppe language AKA immutable Proto-Indo-European: the glottochronological models spanning thousands of years are not valid for the steppe, just as they are not valid for an Anatolian homeland, nor for a Caucasus homeland. The actual cultural-historical early Sredni Stog – Khvalynsk community, formed earlier than ca. 5000 BC, is a thousand years older than the expansion of Khvalynsk with the horse, and some two thousand years older than the expansion of Khvalynsk-Repin/Early Yamna migrants (see here for the latest genetic research).

What lies between the formation of that early Eneolithic cultural-historical community, and what we see in archaeology and genetics in Middle and Late Eneolithic steppe cultures, is the radical differentiation of western (Ukraine Eneolithic, mainly forest-steppe) and eastern (Samara and Khvalynsk/Repin, mainly steppe) cultures and peoples, i.e. precisely the period of differentiation of an eastern, Proto-Indo-Hittite-speaking early Khvalynsk community (that expanded with the horse and horse-related symbols) from a western, probably Early Proto-Uralic speaking community of the North Pontic forest-steppe cultural area.

NOTE. I am not against a Neolithic ‘steppe’ language. But this steppe language was spoken before and/or during the first Neolithisation wave, and should be associated with Indo-Uralic. If there was no Indo-Uralic language, then some communities would have developed Early Proto-Indo-European and Early Proto-Uralic side by side, in close contact to allow for dozens of loanwords or wanderwords to be dated to this period (where, simplistically, PIH *H corresponds to EPU *k, with some exceptions).

steppe-forest-change
Map of a) steppe – forest-steppe border during the Eneolithic in the Pontic-Caspian region and b) the border today, showing a more limited steppe zone in the North Pontic area (reason for the specific ways of expansion of horse-related cultures and horse-related nomadic pastoralism during the Eneolithic).

The convergence that we see in PCA and Admixture of Yamna and the earliest Baltic LN / Corded Ware ‘outlier’ samples (if not directly related exogamy of some Baltic LN/CWC groups with Yamna migrants, e.g. those along the Prut), must be traced back to the period of genetic drift that began precisely with these Khvalynsk-Novodanilovka expansions, also closely associated with populations of the Caucasus, thus bringing North Pontic forest-steppe cultures (probably behind Proto-Corded Ware peoples) nearer to Khvalynsk, and both by extension to Yamna.

We have seen this problem arise in Bell Beaker samples expanding all over Europe, turning from a fully Yamnaya-like population to something else entirely in different regions, from more EEF-like to more CWC-like, sharing one common trait: Y-DNA. We are seeing the same happen with Balkan groups and Mycenaeans, with Old Hittites, and with steppe MLBA from Andronovo peoples expanding over Central and South Asia, and we know that patrilineal clans and thus Y-chromosome bottlenecks were common after Neolithisation, especially with nomadic pastoralist steppe clans (and probably also with many previous population expansions).

Steppe Eneolithic peoples were thus no different to other previous and posterior expanding groups, and ancestry is going to be similar for people living in neighbouring regions, so Y-DNA will remain the essential tool to distinguish different peoples (see here a summary of Proto-Indo-Europeans expanding R1b-L23).

We are nevertheless still seeing “R1b zombies” (a quite appropriate name I read on Anthrogenica) still arguing for a Western European origin of R1b-L23 based on EEF-like ancestry and few steppe-related contribution found in Iberian Bell Beakers (read what David Reich has to say on this question); and “OIT zombies” still arguing for IVC representing Proto-Indo-European, based on Iran_N ancestry and the minimal steppe ancestry-related impact on certain ancient Asian cultures, now partly helped by “Caucasus homeland zombies” with the new PIE=CHG model; apart from many other pet theory zombies rising occasionally from their graves here and there. Let’s hope that this virus of the undead theories does not spread too strongly to the R1a-Indo-European association, when the official data on Khvalynsk, West Yamna, and Yamna Hungary come out and show that they were dominated by R1b-L23 lineages.

Because we need to explore in detail the continuation of Khvalynsk-related (potential Proto-Anatolian) cultures in the Lower Danube and the Balkans, e.g. from Cernavoda I to Cernavoda III, then maybe to Ezero, and then to Troy; as well as the specific areas of Late Indo-European expansions associated with Early Yamna settlers turning into Bell Beakers, Balkan EBA, and Steppe MLBA-associated cultures. There is a lot of work to do on proper definition of Bronze Age cultures and their potential dialects, as well as convergence and divergence trends, and not only of Indo-European, but also of Uralic-speaking communities derived from Corded Ware cultures.

If we let the narratives of the 2000s in Genetics (in combination with the 1960s in Archaeology) dominate the conversation, then a lot of time will be absurdly lost until reality imposes itself. And it will.

EDIT (2 JUL 2018): Some sentences corrected, and some information added to the original post.

Related

Climatic conditions in the Cis-Ural Steppe region and the Repin culture

cis-ural-region

New paper (behind paywall) Climate and Vegetation Changes over the Past 7000 Years in the Cis-Ural Steppe, by Khokhlova, Morgunova, Khokhlov, and Gol’eva, Eurasian Soil Sc. (2018) 51: 506.

Abstract (emphasis mine):

A multilayered archaeological site Turganik Settlement in the valley of the Tok River in the Cis- Ural steppe (Orenburg oblast) was examined with the use of paleopedological and microbiomorph methods. Ancient people inhabited this area in the Latest Neolithic (Eneolithic) (5th millennium BC) and Early Bronze (4th millennium BC) ages. It was found that cultural layers dating back to the Atlantic period of the Holocene had been formed under conditions of a predominance of grassy–forb vegetation with a small portion of tree species and dry climate; the ancient settlement was not affected by floods and was suitable for permanent living. It is probable that soils of the chestnut type with salinization and solonetzic features were developed in that time. The final stages of the accumulation of cultural layers were marked by strong shortterm floods, whose sediments partly masked the features of the previous long arid epoch. The highest degree of aridity was at the end of the Atlantic period. In the Subboreal and Subatlantic periods, soils of the meadowchernozemic type were formed; the spore–pollen spectra of these periods are characterized by a higher portion of tree species and by the presence of phytoliths of meadow grasses. The climatic conditions were generally colder and more humid, though some short-term aridization stages could take place. Some of these stages are recorded in the thickness of the studied sediments.

Interesting excerpts:

Paleosols buried under archaeological monuments of different periods represent a valuable archieve of information about the paleoenvironment. Most of the works in the field of archaeological pedology deal with earthen burial sites and kurgans [2, 5, 7, 9, 11]. Paleosols buried under the kurgans present us the paleoenvironmental records of a relatively short time before they were buried under the kurgan bodies. The study of kurgans created in different times makes it possible to characterize paleosol sequences storing information on longer periods of time in the second half of the Holocene. However, groups of kurgans that were consecutively created during the entire time span of kurgan construction, beginning from about 6000 yrs BC to the Early Medieval epoch, are few in number [8, 18, 22]. Even for such groups of kurgans, there are considerable time intervals that cannot be characterized because kurgans were not constructed during them. Hence, it is impossible to study the soils buried during these intervals. To reconstruct paleoenvironmental conditions for them, certain interpolation is required [29].

At the same time, there are archaeological sites — ancient settlements — the material of which gradually accumulated during very long time, e.g., beginning from the Middle Holocene to the present. Though such objects may also contain “missing layers” for certain periods, when the processes of denudation, erosion, or deflation predominated over sediment accumulation, they represent an almost continuous record on information about the paleoenvironmental conditions from the beginning of their functioning [25]. The most valuable among such sites are those that retain information on those periods of the Holocene that cannot be characterized on the basis of available data on the paleosols buried under the kurgans. In particular, this is the Atlantic period (7500–5000 yrs ago), because the construction of kurgans began only in the second half of this period (within the Volga–Ural interfluve [14]), and such ancient kurgans are rarely found. We studied such an archaeological site in the valley of the Tok River in the Cis-Ural steppe zone in Orenburg oblast. This site is known as the ancient Turganik settlement.

We studied different layers of the Turganik settlement with the use of a set of methods in order reconstruct the paleoenvironmental conditions (climate and vegetation) for the entire period of the accumulation of these sediments.

Thus, the Atlantic period of the Holocene in the Cis-Uralian steppe was characterized by dry climatic conditions with the driest stage during the Early Bronze Age (the Early Yamnaya culture of the middle of the fourth millennium BC). The Subboreal and Subatlantic periods were generally colder and wetter, though they also included short-term aridization phases, some of which were recorded in the sediment thickness.

This site is at the core of the interaction of Samara, Khvalynsk, and Repin cultures during the Eneolithic.

You can read more about it and the nature of Repin described by Morgunova (in favour of Gimbutas’ model), as combining traditions from Eneolithic steppe cultures from Khvalynsk to Sredni Stog, e.g. in Pottery from the Volga area in the Samara and South Urals region from Eneolithic to Early Bronze Age (2015).

repin-khvalynsk
Eneolithic settlements (1–5, 7, 10–16, 20, 22–43, 48, 50), burial grounds (6, 8–9, 17–19, 21, 47, 49) and kurgans (44–46) of the steppe Ural-Volga region: 1 Ivanovka; 2 Turganik; 3 Kuzminki; 4 Mullino; 5 Davlekanovo; 6 Sjezheye (burial ground); 7 Vilovatoe; 8 Ivanovka; 9 Krivoluchye; 10–13 LebjazhinkaI-III-IV-V; 14 Gundorovka; 15–16 Bol. Rakovka I-II; 17–18 Khvalunsk I-II; 19 Lipoviy Ovrag; 20 Alekseevka; 21 Khlopkovskiy; 22 Kuznetsovo I; 23 Ozinki II; 24 Altata; 25 Monakhov I; 26 Oroshaemoe; 27 Rezvoe; 28 Varpholomeevka; 29 Vetelki; 30 Pshenichnoe; 31 Kumuska; 32 Inyasovo; 33 Shapkino VI; 34 Russkoe Truevo I; 35 Tsaritsa I-II; 36 Kamenka I; 37 Kurpezhe-Molla; 38 Istay; 39 Isekiy; 40 Koshalak; 41 Kara-Khuduk; 42 Kair-Shak VI; 43 Kombakte; 44 Berezhnovka I-II; 45 Rovnoe; 46 Politotdelskoe; 47 burial near s. Pushkino; 48 Elshanka; 49 Novoorsk; 50 Khutor Repin.

The migration model of Anthony (2007, 2015), who collaborated with this group, is a more precise description of how peoples from the east of the Don River (mainly Khvalynsk/Repin cultures) migrated to develop a greater Yamna community, with Repin-type material culture expanding east of the Urals (into Afanasevo) and west of the Don River (into previous Sredni Stog/Kvitjana territory) – which I followed for the Indo-European demic diffusion model (we have recent samples of other potential Khvalynsk/Repin-related migrations).

NOTE. I usually refer to this Khvalynsk/Repin migration in genomics as of ‘Khvalynsk migrants’, for simplicity purposes, given that the few samples we have are from Khvalynsk, and that cultural regions east of the Don are difficult to differentiate precisely. However, it remains to be seen if – as I proposed – there are genetic differences between Repin and Khvalynsk groups, especially regarding R1b-L23 subclades – I proposed mainly Z2103 for Khvalynsk, L51 for Repin, a difference which has not been confirmed for the moment in Afanasevo, probably of Pre-Tocharian dialect, an archaic Northern dialect of Late PIE.

Anthony’s model of Khvalynsk/Repin as Yamna forefathers is probably, as we are seeing in Yamna samples, the right interpretation of peoples behind pots, compared to Gimbutas’ general idea of expanding kurgans of the 1970s.

On the other hand, the alternative Russian school version of Yamna developing from a heterogeneous community of Khvalynsk-Sredni Stog-Lower Danube cultures is probably by now to be fully dismissed, in archaeology (as Morgunova says) as in genetics.

Related:

Steppe and Caucasus Eneolithic: the new keystones of the EHG-CHG-ANE ancestry in steppe groups

indo-uralic-ehg-chg-ane-ancestry

Some interesting excerpts from Wang et al. (2018):

An interesting observation is that steppe zone individuals directly north of the Caucasus (Eneolithic Samara and Eneolithic steppe) had initially not received any gene flow from Anatolian farmers. Instead, the ancestry profile in Eneolithic steppe individuals shows an even mixture of EHG and CHG ancestry, which argues for an effective cultural and genetic border between the contemporaneous Eneolithic populations in the North Caucasus, notably Steppe and Caucasus. Due to the temporal limitations of our dataset, we currently cannot determine whether this ancestry is stemming from an existing natural genetic gradient running from EHG far to the north to CHG/Iran in the south or whether this is the result of farmers with Iranian farmer/ CHG-related ancestry reaching the steppe zone independent of and prior to a stream of Anatolian farmer-like ancestry, where they mixed with local hunter-gatherers that carried only EHG ancestry.

PCA-caucasus-khvalynsk-sredni-stog
Image modified from Wang et al. (2018). Samples projected in PCA of 84 modern-day West Eurasian populations (open symbols). Previously known clusters have been marked and referenced. An Eastern European (blue) and a Caucasus (brown) ‘clouds’ have been drawn in dotted circles, leaving Pontic-Caspian steppe and derived groups between them.See the original file here.

Concerning the influences from the south, our oldest dates from the immediate Maykop predecessors Darkveti-Meshoko (Eneolithic Caucasus) indicate that the Caucasus genetic profile was present north of the range ~6500 BP, 4500 calBCE. This is in accordance with the Neolithization of the Caucasus, which had started in the flood plains of the great rivers in the South Caucasus in the 6th millennium BCE from where it spread to the West and Northwest Caucasus during the 5th millennium BCE9, 49. It remains unclear whether the local CHG ancestry profile (represented by Late Upper Palaeolithic/Mesolithic individuals from Kotias Klde and Satsurblia in today’s Georgia) was also present in the North Caucasus region before the Neolithic. However, if we take the Caucasus hunter-gatherer individuals from Georgia as a local baseline and the oldest Eneolithic Caucasus individuals from our transect as a proxy for the local Late Neolithic ancestry, we notice a substantial increase in Anatolian farmer-related ancestry. This in all likelihood is linked to the process of Neolithization, which also brought this type of ancestry to Europe. As a consequence, it is possible that Neolithic groups could have reached the northern flanks of the Caucasus earlier50 (Supplementary Information 1) and in contact with local hunter gatherers facilitated the exploration of the steppe environment for pastoralist economies. Hence, additional sampling from older individuals is needed to fill this temporal and spatial gap.

The newest paper of the Reich/Jena group has brought samples (probably) much nearer to the actual CHG and ANE contribution seen in Eneolithic steppe peoples than the previously available Kotias Klde, Satsurblia, Afontova Gora 3, or Mal’ta.

It is impossible to say without direct access to the samples, but it is very likely that we will soon be able to break down different gross contributions from groups similar to these Steppe/Caucasus Neolithic ancestral groups into the diverse Eneolithic cultures of the Pontic-Caspian steppe, and thus trace more precisely each of these cultures to their genetic (and thus ethnolinguistic) heirs.

qpgraph-eneolithic-steppe
Admixture Graph modelling of the population history of the Caucasus region. We started with a skeleton tree without admixture including Mbuti, Loschbour and MA1. We grafted onto this EHG, CHG, Globular_Amphora, Eneolithic_steppe, Maykop, and Yamnaya_Caucasus, adding them consecutively to all possible edges in the tree and retaining only graph solutions that provided no differences of |Z|>3 between fitted and estimated statistics. The worst match is |Z|=2.824 for this graph. We note that the maximum discrepancy is f4(MA1, Maykop; EHG, Eneolithic_steppe) = -3.369 if we do not add the 4% EHG ancestry to Maykop. Drifts along edges are multiplied by 1000 and dashed lines represent admixture.”

Some more representative samples from Eneolithic steppe, steppe-forest and forest zone cultures of Eastern Europe will probably help with the fine-scale structure of different Chalcolithic groups, especially the homeland of early Corded Ware groups.

These new samples seem another good reason (like the Botai and R1b-M73) to rethink the role of (what I assumed were) different westward Mesolithic Eurasian waves of expansion influencing the formation of an Indo-Uralic and Indo-European community in Eastern Europe, and return to the simpler idea of local contributions from North Caucasus and steppe peoples absorbed by expanding EHG-like groups.

Related:

Post-Neolithic Y-chromosome bottleneck explained by cultural hitchhiking and competition between patrilineal clans

Open access study Cultural hitchhiking and competition between patrilineal kin groups explain the post-Neolithic Y-chromosome bottleneck, by Zeng, Aw, and Feldman, Nature Communications (2018).

Abstract (emphasis mine):

In human populations, changes in genetic variation are driven not only by genetic processes, but can also arise from cultural or social changes. An abrupt population bottleneck specific to human males has been inferred across several Old World (Africa, Europe, Asia) populations 5000–7000 BP. Here, bringing together anthropological theory, recent population genomic studies and mathematical models, we propose a sociocultural hypothesis, involving the formation of patrilineal kin groups and intergroup competition among these groups. Our analysis shows that this sociocultural hypothesis can explain the inference of a population bottleneck. We also show that our hypothesis is consistent with current findings from the archaeogenetics of Old World Eurasia, and is important for conceptions of cultural and social evolution in prehistory.

Relevant excerpts:

y-dna-bottleneck
Tree of Y-chromosome genotypes from samples found among cultures with hunter-gatherer subsistence, and agropastoralist subsistence. The blue background represents hunter-gatherer subsistence while the green background represents agropastoralist subsistence. Letters in red circles match individuals from sites with their archaeological context. Note that R1b-P321 is synonymous with R1b-S116. Adapted from Figs. 3, 4, 5 and 6 of Kivisild67, with addition of information from Olalde et al.64. The vertical axis represents time; the position of branch points represent the ages of branch-defining mutations, with nomenclature and age from yfull (https://www.yfull.com/tree/)

Our hypothesis explains the bottleneck as a consequence of intergroup competition between patrilineal kin groups, which caused cultural hitchhiking between Y-chromosomes and cultural groups and reduction in Y-chromosomal diversity. Competition between demes can dramatically reduce genetic diversity within a population1, especially if the population is structured such that variation is greater between demes than within demes. Culturally transmitted kinship ideals and norms can cause homophilous sorting and limit interdemic gene flow, creating homogeneous demes that differ strongly from one another. Patrilineal corporate kin groups, with coresiding male group members descending from a common male ancestor, would produce such an effect on Y-chromosomes only, as patrilineal corporate kin groups generally coexist with female exogamy40, which would homogenize the mitochondrial gene pools of different groups41,42.

With intergroup competition between patrilineal corporate kin groups, two mechanisms would operate to reduce Y-chromosomal diversity. First, patrilineal corporate kin groups produce high levels of Y-chromosomal homogeneity within each social group due to common descent, as well as high levels of between-group variation. Second, the presence of such groups results in violent intergroup competition preferentially taking place between members of male descent groups, instead of between unrelated individuals. Casualties from intergroup competition then tend to cluster among related males, and group extinction is effectively the extinction of lineages.

There is evidence that other analogous situations involving gene-culture hitchhiking in culturally-defined social groups may have affected genetic diversity. Central Asian pastoralists, who are organized into patriclans, have high levels of intergroup competition and demonstrate ethnolinguistic and population-genetic turnover down into the historical period59. They also have a markedly lower diversity in Y-chromosomal lineages than nearby agriculturalists42,60. In fact, Central Asians are the only population whose male effective population size has not recovered from the post-Neolithic bottleneck; it remains disproportionately reduced, compared to female estimates using mtDNA4. Central Asians are also the only population to have star-shaped expansions of Y-chromosomes within the historical period, which may be due to competitive processes that led to the disproportionate political success of certain patrilineal clans60.

The simulation offers an interesting graphic. I had been thinking for some time about developing an interactive image with waves of expansion showing how only few haplogroups expand and thus their variability is reduced in successive migration waves, because a lot of people seemed not to be willing to accept this:

y-dna-bottleneck-simulation
Schematic of the steps in the simulation, according to the order described in the algorithm. a (i) Patrilineal (PT) starting conditions, where cultural groups strictly determine haplogroup type. a (ii) The non-patrilineal (NPT) condition where they are perfectly uncorrelated. b The killing step, with a more (PT) and less (NPT) patrilineal starting condition. The number of deaths in each group is inversely related to group size. The blue cultural group goes extinct in both cases. This causes the haplogroup represented by the diamonds to go extinct in PT, but no haplogroup extinction occurs in NPT. c The mutation step, where a small number of individuals in the largest haplogroup change their haplogroup. d The regeneration step, where (i) is a replica of (b) PT (iii), and (d) (ii) shows how the original number of individuals before the killing step is restored by proportionally increasing the number of individuals in all cells. e Group fission step. Where an empty row occurs, the largest cultural group splits, and half the individuals form a new cultural group in the empty row. The step in which we remove cultural groups that are too small—between (c, d) (see Methods)—is not shown

You only have to imagine this process happening in many successive waves of expansion (external as well as internal to each culture) since the first Neolithic expansions in the steppe in the late-6th millennium BC, even before the formation of the Khvalynsk-Sredni Stog cultural-historical community, to understand what happened in the next thousands of years with evolving patrilineal clans and their distinct cultures.

The whole paper is an interesting read. It’s great to see sociology and genetics finally catch up and interact to develop more complex anthropological hypotheses.

The fact that this paper appears in mid-2018 and geneticists are beginning to discuss this only now when their statistical methods fail to explain the obvious (see David Reich’s recent interview) seems anachronistic, though, because all this was quite clear already in 2015 – at least for those who were looking for mainstream Yamna – Bell Beaker connections, instead of inventing new migration pathways to justify the results of certain statistical analyses

Anyway, better late than never.

Also, they use YFull estimates, which vindicates my use of them in the Indo-European demic diffusion model (2017). On the other hand, their use of these estimates right now in 2018 for R1a-M417 and R1b-M269 – when we know of a R1a-Z93 case much older than YFull’s estimated 5,000 YBP for this subclade, and possibly for R1b-L23, too, is the biggest pitfall in their temporal assessment, although the bottlenecks seen in Chalcolithic expansions seem to have indeed began during the Mesolithic-Neolithic transition in the steppe.

So, say goodbye (if you haven’t already) to dat fantasy ‘steppe people’ of mixed R1a/R1b descent cooperating with the same mixed steppe language, all represented by the Yamnaya™ ancestral component, and say hello to distinct, competing ethnolinguistic steppe groups during the Neolithic.

Related:

East Bell Beakers, an in situ admixture of Yamna settlers and GAC-like groups in Hungary

indo-european-yamnaya-corded-ware

I wanted to repeat what I said last week in two different posts (see on the new Caucasus and Yamna Hungary samples, and on local groups in contact with Yamna settlers).

We already knew that expanding East Bell Beakers had received influence from a population similar to the available Globular Amphorae culture samples.

  1. Without Yamna settlers, but with Yamna Ukraine and East Bell Beaker samples, including an admixed Yamna Bulgaria sample (from Olalde & Mathieson 2017, and then with their Nature 2018 papers), the most likely interpretation was that Yamna settlers had received GAC ancestry probably during their migration through the Balkans, before turning into East Bell Beakers. However, some comments still supported that it was Corded Ware migrants the ones behind the formation of East Bell Beakers. I couldn’t understand it.
  2. Now we have (with Wang et al. 2018) Yamna settlers (identical to other Yamna groups and Afanasevo migrants) and GAC-like peoples coexisting with them in Hungary, with a Late Chalcolithic Yamna sample from Hungary showing a greater contribution from GAC. However, I still read discussions on Yamna settlers receiving GAC admixture from Corded Ware in Eastern Europe, from GAC in the Dnieper-Dniester area, in Budzhak/Usatovo, etc. I can’t understand this, either.
  3. I will post here the data we have, with the simplest maps and images showing the simplest possible model. No more long paragraphs.

    NOTE. All this data does not mean that this model is certain, especially because we don’t have direct access to the samples. But it is the simplest and most likely one. Sometimes 2+2=4. Even if it turns out later to be false.

    EDIT (30 MAY 2018): In fact, as I commented in the first post about these samples, there is a Yamna LCA/EBA sample probably from Late Yamna (in the North Pontic steppe, west of the Catacomb culture), which shows GAC-like contribution. However, this admixture is lesser than that of Hungary LCA/EBA1 sample, and both Yamna groups (Hungary and steppe) were probably already more sedentary, which also supports different contributions from nearby local GAC-like groups to each region, rather than maintained long-range internal genetic contributions from a single source near the steppe…

    indo-european-uralic-migrations-yamna-gac
    Yamna migrants ca. 3300-2600. Most likely site of admixture with GAC circled in red.
    yamna_bell_beaker
    Yamna – Bell Beaker migration according to Heyd (2007, 2012). Most likely site of admixture with GAC is marked by the evolution of Blue to Red color.
    PCA-yamna-hungary
    PCA results. Samples from Yamna Hungary are surrounded by red circles, GAC-like Hungarian groups surrounded by light brown (see below for ADMIXTURE data) Notice the most likely Yamna Hungary sample with GAC admixture clustering closely to CWC Esperstedt outlier, and thus to some East Bell Beaker samples. (d) shows these projected onto a PCA of 84 modern-day West Eurasian populations (open symbols).
    gac-like-hungary-yamnaya
    Modified image, with red rectangles surrounding (unreleased) Hungarian samples from Yamna and GAC-like groups. (c) ADMIXTURE results of relevant prehistoric individuals mentioned in the text (filled symbols)
    yamnaya-hungary-lca-eba
    Modified image, with red rectangles surrounding (unreleased) Yamna samples Notice greater GAC contribution to late Yamna Hungary sample. Modelling results for the Steppe and Caucasus cluster. Admixture proportions based on (temporally and geographically) distal and proximal models, showing additional Anatolian farmer-related ancestry in Steppe groups as well as additional gene flow from the south in some of the Steppe groups as well as the Caucasus groups
    yamnaya-hungary-globular-amphora
    Modified table from Wang et al. (2018) Supplementary materials (in bold, Yamna and related samples; in red, newly reported samples). Notice greater GAC contribution to late Yamna Hungary sample. “Supplementary Table 18. P values of rank=1 and admixture coefficients of modelling the Steppe ancestry populations as a two-way admixture of the Eneolithic_steppe and Globular_Amphora using 14 outgroups. Left populations: Steppe cluster, Eneolithic_steppe, Globular Amphora Right populations: Mbuti.DG, Ust_Ishim.DG, Kostenki14, MA1, Han.DG, Papuan.DG, Onge.DG, Villabruna, Vestonice16, ElMiron, Ethiopia_4500BP.SG, Karitiana.DG, Natufian, Iran_Ganj_Dareh_Neolithic.”

    The CWC outlier from Esperstedt

    I already said that my initial interpretation of the Esperstedt outlier, dated ca. 2430 BC, as due to a late contribution directly from the steppe (i.e. from long-range contacts between late Corded Ware groups from Europe and late groups from the steppe) was probably wrong, seeing how (in Olalde et al. 2017) early East Bell Beaker samples from Hungary and Central Europe clustered closely to this individual.

    Now we see that fully ‘Yamnaya-like’ Yamna settlers lived in Hungary probably for two or three centuries ca. 2900-2600 BC, and the absorption of known (or unknown) Yamna vanguard groups found up to Saxony-Anhalt before 2600 BC would be enough to justify the genomic findings of this individual.

    An outlier it is, then. But probably from admixture with nearby Yamna-like people.

    olalde_pca
    Image modified by me, from Olalde et al. (2017). PCA of 999 Eurasian individuals. Marked is the Espersted Outlier.

    Related:

The R1b-L23/Late PIE expansions, and the ‘R1a – Indo-European’ association

indo-european-yamnaya-corded-ware

I wrote a series of posts at the end of 2017 / beginning of 2018, to answer the wrong assumptions I could read in forums and blogs since 2015.

I decided not to publish them then, seeing how many successive papers were confirming my Indo-European demic diffusion model in a (surprisingly) clear-cut way.

Nevertheless, because I keep reading the same comments no matter what gets published, even in mid-2018 – the latest ones in our Facebook page (“was haplogroup X Indo-European?”), and in this very blog (“I see it very difficult to link Bell Beaker with Balto-Slavic, when now Balto-Slavic people are strikingly R1a-dominated”); and because I see even more misunderstandings and personal attacks, I have decided to publish them.

This way I will be able to explain my “R1b-L23/Proto-Indo-Europeans” theory with simplistic maps (however badly I hate such maps when I find them on Google searches), and I will also have a page to redirect those who don’t want to dismiss the “R1a – Indo-European association”, instead of answering comments about this question each time they pop up…

Here you have the links to the posts – and also on the menu above (there is a lot of rambling, because they are from a period of less clear data on Yamna and Corded Ware; today I would have never written such long discussions, they are mostly unnecessary):

  1. Haplogroup is not language, but R1b-L23 expansion was associated with Proto-Indo-Europeans
  2. The history of the simplistic ‘haplogroup R1a — Indo-European’ association
  3. Tips for dialogue with those supporting the R1a/Indo-European association

Related:

Copenhagen group: Germanic and Balto-Slavic from Bell Beaker; Indo-Anatolian homeland in the Caucasus

invasion-from-the-steppe-yamnaya

Article of general knowledge in Der Spiegel, Invasion from the Steppe, with comments from Willerslev and Kristiansen, appeared roughly at the same time as the Damgaard et al. Nature (2018) and Science (2018) papers were published.

NOTE. You can read the article (in German) from Kristiansen’s Academia.edu account.

Excerpts translated from German (emphasis mine):

On the Y-DNA data

Particularly striking is the genetic signature from the steppe on the Y chromosome. From this the researchers conclude that the majority of migrants were males. Kristian Kristiansen, chief archaeologist in the Willerslev team, also has an idea of ​​how this could be explained: “Maybe it’s a rite of initiation, as it was spread among the steppe peoples,” he says.

The younger sons of the Yamnaya herders, who were excluded from the succession, had to seek their fortune on their own. As part of a solemn ritual, they threw themselves to wolves’ skins and then swarmed in warlike gangs to buy their own herds by cattle-stealing.

(…)

An ally that they seem to have brought from their homeland may also have contributed to the genetic success of the steppe people: Yersinia pestis, the plague bacterium. Its genes were found by researchers from the Max Planck Institute in Jena – and apparently it emerged exactly at the same time as the Yamnaya thrust began.

About the Hittites

(…) And yet now, where Asia and Europe meet geographically, there is no trace of the Yamnaya genes. The wander-loving people from the Pontic-Caspian steppe apparently found neither the way across the Balkans nor through the Caucasus mountains.

Now the researchers are puzzled: How can it be that a language goes on a walk, without the accompanying speakers coming along? Is it possible that the Indo-European seeped into Anatolia, much like the English language spread today without the need for Englishmen?

Archaeologist Kristiansen does not believe it. The researchers would find it hard to reconsider their theories, he says: “Especially the first chapter of the story has to be rewritten.”

He suspects that there was a predecessor of the Yamnaya culture, in which a kind of Proto-Proto-Indo-European was spoken. And he also has a suspicion, where this people could have drifted around: The Caucasus, says Kristiansen, was their homeland. But that remains unproven: “There’s another hole left,” he admits.

invasion-yamnaya-steppe
Spread of Indo-European languages

About the Botai

The study of [the Botai] genome revealed that it was genetically radically different from the members of the Yamnaya culture. The Botai, it seems, consistently avoided any contact with their neighbors – even though they must have crossed the territory of the Botai on their migratory waves.

Willerslev assumes that the art of keeping horses from the Yamnaya steppe nomads was adopted from these peoples, and then they developed it further. At some point, the Botai could then have itself become doomed by its groundbreaking innovation: While the descendants of the Yamnaya spread over half of Eurasia, the Botai disappeared without leaving a trace.

Even more interesting than the few words that set the Copenhagen group’s views for future papers (such as the expected Maykop samples with EHG ancestry) is the artistic sketch of the Indo-European migrations, probably advised by the group.

A simple map does not mean that all members of the Danish workgroup have changed their view completely, but I would say it is a great improvement over the previous “arrows of migration” (see here), and it is especially important that they show a more realistic picture of ancient migrations to general readers.

NOTE. Especially absurd is the identification of the ‘Celtic’ expansion with the first Bell Beakers in the British Isles (that idea is hold by few, such as Koch and Cunliffe in their “Celtic from the West” series). Also inexact, but not so worrying, are the identification of ‘Germanic’ in Germany/Únětice, or the spread of ‘Baltic’ and ‘Slavic’ directly to East Europe (i.e. I guess Mierzanowice/Nitra -> Trzciniec), which is probably driven by the need to assert a close connection with early Iranians and thus with their satemization trends.

Also, as we know now thanks to Narasimhan et al. (2018), there is no need to support that convoluted west arrow (representing CWC) from West Yamna to Central Europe, and then to East Yamna, since the Proto-Indo-Iranian community – represented by the Steppe MLBA cloud that later expanded Indo-Aryan and Iranian languages – has a more direct connection with the in situ admixture of Poltavka/Abashevo within the Volga-Ural region.

I think we can keep this from the article:

Their results, as well as those of the competition labs at Harvard University and Jena’s Max Planck Institute for the History of Humanity, leave no doubt: Yes, the legendary herdsmen in the Pontic-Caspian steppe really existed. They belonged to the so-called Yamnaya culture, and they spread, as linguists had predicted, in massive migrations towards Central Europe and India – a later triumph for linguists.

This can be added to a recent comment by de Barros Damgaard:

The project has been an extremely enriching and exciting process. We were able to direct many very different academic fields towards a single coherent approach. By asking the right questions, and keeping limitations of the data in mind, contextualizing, nuancing, and keeping dialogues open between scholars of radically different backgrounds and approaches, we have carved out a path for a new field of research. We have already seen too many papers come out in which models produced by geneticists working on their own have been accepted without vital input from other fields, and, at the other extreme, seen archaeologists opposing new studies built on archaeogenetic data, due to a lack of transparency between the fields.

Data on ancient DNA is astonishing for its ability to provide a fine-grained image of early human mobility, but it does stand on the shoulders of decades of work by scholars in other fields, from the time of excavation of human skeletons to interpreting the cultural, linguistic origins of the samples. This is how cold statistics are turned into history.

Related:

Immigration and transhumance in the Early Bronze Age Carpathian Basin

Interesting excerpts about local Hungarian groups that had close contacts with Yamna settlers in the Carpathian Basin, from the paper Immigration and transhumance in the Early Bronze Age Carpathian Basin: the occupants of a kurgan, by Gerling, Bánffy, Dani, Köhler, Kulcsár, Pike, Szeverényi & Heyd, Antiquity (2012) 86(334):1097-1111.

The most interesting of the local people is the occupant of grave 12, which is the earliest grave in the kurgan and the main statistical range of its radiocarbon date clearly predates the arrival of the western Yamnaya groups c. 3000 BC. This is also confirmed by the burial rite, which is not typical for the Yamnaya (Dani 2011: 29–33; Heyd in press), although some heterogeneity may apply in Yamnaya communities too. The migrant group, graves nos. 4, 7, 9 and 11, all occupy late stratigraphic positions in the mound, and have radiocarbon dates in the second quarter of the third millennium BC. It is also noteworthy that they are all adult or mature men. The contextual data, their physical distribution over the space of the whole kurgan, and the variety of burial practices, indicate several generations of burials. The cultural attributes of this group are summarised in Figure 5. Overall, their closest match lies in the Livezile group from the eastern and southern Apuseni Mountains, which is also the likely place of origin of the buried persons.

yamna-settlements-hungary
Cultural geography of the Carpathian Basin in the first half of the third millennium BC (in black: archaeological cultures and groups dating roughly to the first quarter; in red: those dating to the second quarter). Indicated also are regions and sites mentioned in the text.

The key question is, what cultural process could be responsible for attracting these men from their homeland to the Great Hungarian Plain, over several generations? Their sex and age uniformity indicate they are a social sub-set within a larger group, implying that only a portion of their society was on the move. Exogamy can probably be excluded, since one would expect more women than men to move in prehistoric times; not to mention the distance of more than 200km between the places of potential origin and burial.

One hypothesis would see these men involved in the exchange of goods, with long-term relations between the mountain and steppe communities. Normally living in, or next to, the Apuseni, these men would journey for weeks into the plain, returning to the same places and people over many decades. Ethnographic examples of such travels to exchange objects and ideas, and perhaps people, are numerous (e.g. Helms 1988). However, the child’s (grave 7a) local isotopic signature would remain unexplained, and one has to wonder for how many generations an exchange continues for four men to die near the Őrhalom.

A second hypothesis is essentially an economic model of transhumance, with livestock passing the winter and spring in the milder regions of the Great Hungarian Plain, and returning to higher pastures in the warmer months (Arnold & Greenfield 2006). Such systems can endure for centuries, provided the social relations underpinning them are stable. This has the advantage of accounting for relatively long periods of time spent away from home, as herdsmen guarded their animals, and perhaps some women and their children came too, which would account for the child’s presence, and the pottery relations of the Livezile group. Furthermore, regular visits to a region would increase the likelihood of Livezile transhumant herders becoming integrated locally. The second quarter of the third millennium BC was a period when Yamnaya ideology, and thus its internal coherence, might have already diminished. This would likely have resulted in a weakened grip by Yamnaya people on pastures and territory, consequently allowing Livezile herders, and potentially others, to step in and take over locally, perhaps first on a seasonal basis and then permanently.

On West Yamna settlers in Hungary

yamnaya-hungary-globular-amphora
Modified table from Wang et al. (2018) Supplementary materials (in bold, Yamna and related samples; in red, newly reported samples). “Supplementary Table 18. P values of rank=1 and admixture coefficients of modelling the Steppe ancestry populations as a two-way admixture of the Eneolithic_steppe and Globular_Amphora using 14 outgroups. Left populations: Steppe cluster, Eneolithic_steppe, Globular Amphora Right populations: Mbuti.DG, Ust_Ishim.DG, Kostenki14, MA1, Han.DG, Papuan.DG, Onge.DG, Villabruna, Vestonice16, ElMiron, Ethiopia_4500BP.SG, Karitiana.DG, Natufian, Iran_Ganj_Dareh_Neolithic.”

By disclosing very interesting information on (yet unpublished) Yamna samples from Hungary, the latest preprint from the Reich Lab has rendered irrelevant – in a rather surprising turn of events – (what I expected would be) future discussions on West Yamna settlers potentially sharing a similar ancestry with Baltic Late Neolithic / Corded Ware settlers (see here for more details).

Interesting excerpts regarding the tight cluster formed by all Yamna samples:

Individuals from the North Caucasian steppe associated with the Yamnaya cultural formation (5300-4400 BP, 3300-2400 calBCE) appear genetically almost identical to previously reported Yamnaya individuals from Kalmykia20 immediately to the north, the middle Volga region19, 27, Ukraine and Hungary, and to other Bronze Age individuals from the Eurasian steppes who share the characteristic ‘steppe ancestry’ profile as a mixture of EHG and CHG/Iranian ancestry23, 28. These individuals form a tight cluster in PCA space (Figure 2) and can be shown formally to be a mixture by significantly negative admixture f3-statistics of the form f3(EHG, CHG; target) (Supplementary Fig. 3).

Using qpAdm with Globular Amphora as a proximate surrogate population (assuming that a related group was the source of the Anatolian farmer-related ancestry), we estimated the contribution of Anatolian farmer-related ancestry into Yamnaya and other steppe groups. We find that Yamnaya individuals from the Volga region (Yamnaya Samara) have 13.2±2.7% and Yamnaya individuals in Hungary 17.1±4.1% Anatolian farmer-related ancestry (Fig.4; Supplementary Table 18)– statistically indistinguishable proportions.

yamna_bell_beaker
Yamna – Bell Beaker migration according to Heyd (2007, 2012)

Before this paper, we had the solidest anthropological models backed by Y-DNA against conflicting data from certain statistical tools applied to a few samples (which some used to contradict what was mainstream in Academia).

NOTE. I have discussed this extensively in this blog, and more than once. See for example my posts on R1a speaking IE (July 2017), on the Eneolithic Ukraine sample (September 2017), or on the “Yamnaya ancestral component” (November 2017).

Today, we have everything – including statistical tools – showing a genetically homogeneous, Late PIE-speaking late Khvalynsk/Yamna community expanding into its known branches, confirming what was described using traditional anthropological disciplines:

  • Late Khvalynsk expanding into Afanasevo ca. 3300-3000 BC with an archaic Late PIE dialect, which was attested much later as Tocharian;
  • East Yamna/Poltavka admixing with Uralic-speaking Abashevo migrants probably ca. 2600-2100 BC to form Proto-Indo-Iranian-speaking Sintashta-Petrovka and Potapovka;
  • and now also Yamna settlers: those in Hungary admixing (probably ca. 2800-2500 BC) with the local population to form North-West Indo-European-speaking East Bell Beakers; those from the Balkans forming other IE-speaking Balkan cultures, including the peoples that admixed in Greece, as seen in Mycenaeans.

If Volker Heyd is right with this and other papers – and he has been right until now in his predictions regarding Yamna, Bell Beaker, and Corded Ware cultures – , the change in ancestry will probably begin to be noticed in Yamna samples from Hungary and the Lower Danube during the second quarter of the 3rd millennium, a period defined by the addition of a more fashionable western Proto-Bell Beaker package to the fading traditional Yamna cultural package.

EDIT (19 MAY 2018): I corrected some sentences and added interesting information.

Related:

The unique elite Khvalynsk male from a Yekaterinovskiy Cape burial

Recent paper (behind paywall) The Unique Burial of the Ekaterinovsky Cape Early Eneolithic Cemetery in the Middle Volga Region, by Korolev et al. Stratum Plus (2018) Nº2.

Abstract (official, in English):

This is the first time we published the results of a comprehensive study of burial 45 of the eneolithic cemetery called Ekaterinovsky Cape. The burial contains the skeleton of a young man with traumatic injuries of the skull, leg and hand bones of other individuals, skeleton of a young specimen of a domestic goat (Capra hircus) that was abundantly sprinkled with red ocher. Grave goods include three stone scepters of different types, a large item made of horn in the shape of a bird’s head, a stone adze, knife-like plates of quartzite, beads from the flaps of the shells (Unio), marmot cutters, decoration made from a beaver’s tooth. The uniqueness of the burial is determined by the combination of the composition of the grave goods and traces of ritual practices. To conclude, we suggest the buried man could belong to the elite of the society that left this burial ground.

NOTE. About my terminology, Russian has a lenited pronunciation of E in this case, hence the “Ye-” transliteration of the name of the town (and the site) in Google as Yekaterinovka. The “more etymological” transliteration is with “E”, as they use here, although Russians paradoxically use phonetic transliterations of foreign terms. I prefer the lenited transliteration to distinguish the Russian site from other Ekaterinovkas, though.

ekaterinovsky
Schematic view from burial 45. Male of 20-25 years, ca. 4400-4200 BC.

Interesting excerpt (translated from Russian):

Perhaps, we should correlate three very closely related damages [on the skull] with certain rituals, with which scepters could be associated. Each scepter could be a symbolic expression of a part of society, a type of activity, reaching a certain age and social status. This assumption does not seem incredible in combination with other extant, no less impressive, details of the funeral rite. Of great interest is the ornithomorphic rod of the horn. The location of the wand in the head and right half of the breast emphasizes its special significance in ritual practice and in funeral rites. Direct analogies to this product in other burial places of the cemetery are absent, and outside it authors are not known.

NOTE. Although the paper is in Russian and is behind paywall, it is really cheap, and can be easily translated with Google Translate if you can’t read Russian, so – unlike usual papers from the big publishing companies – you could support the journal by paying for it. You can read more about this burial at Pikabu, too. Photos and text in that post are not the same as in the paper, though, so it seems that the author of the text got the information either directly or from another source.

On the genetic data

Here is what I could gather about the report I shared of R1b-L51 lineages in Samara:

1) Yes, the comment at MolGen.org contains a more or less accurate summary of the oral communication actually given. And no, no more interesting data – from a genetic point of view – was presented.

2) What A.A. Khokhlov reported was preliminary genetic information from some samples, and an outside lab shared this information with him.

NOTE. It is well-known that David Anthony, also part of the Samara Valley project, provided the Reich Lab with Khvalynsk and Yamna samples from the region, so it would not be a surprise that these had been in fact assessed by the Reich Lab, too. This is my assumption, though, and I may be wrong.

3) What the report conveys is that “all samples investigated” belonged to R1b-P312 and R1b-U106, so I understand there are in principle more than two samples, whatever Google Translate says.

4) As R. Rocca said in Anthrogenica, the reported R1b1a1a2a1a1c2b2b1a2 (U106 subclade) is exactly the same one reported in Narasimhan et al. (2018) for the sample from the Iron Age site Loebanr 1 (Swat proto-historic graves) ca. 950 BC.

NOTE. That would be another hint at the origin of the preliminary data, together with the timing of the report (January), probably coinciding with the final assessment of samples which appeared in Narasimhan et al. (2018). That would explain the similar weird Y-SNP calls from software yHaplo (as reported by Narasimhan in Twitter). This is all again conjecture, though.

R1b-P312 is not reported in Narasimhan et al. (2018) for any sample (that would be “R1b1a1a2a1b”, following the standard used in their tables). Because the V88 sample in Khvalynsk, as well as other previously known V88 samples, are correctly reported as within the V88 branch, we may be talking about anything in the R1b tree from L754 (xV88) on. Most likely at or beyond the subclade of the Zvejnieki sample of hg R1b1a1 (classified as of R1b1a1a2a1), i.e. from P297 on.

NOTE. Since R1b-Z2103 samples are correctly reported, it is unlikely that the reported samples are from this branch, either.

ekaterinovka-burial-45
Graphic reconstruction from the elite male of grave 45, by R. M. Galeev.

It is possible, then, that we will have haplogroup R1b-M269 or L23 instead of L51, after all, and there would be then no major corrections to be made, either to the estimated dates from McDonald or Yfull (with their current differences), or to my predictions for early and late Khvalynsk, Repin, and Yamna

NOTE. In fact, the appearance of R1b-M269* and/or L23* linked to expanding Khvalynsk could be the perfect end to the resurging theories on Armenian or Western European origin of this haplogroup.

5) The full official genetic data is expected within a year (precise date unknown), so unless someone knows of a related draft in the making (which could publish them earlier), I would keep my expectations low for an official confirmation of the precise subclade any time soon.

NOTE. The best likely proxy for the reported data, if the above assumptions on Y-SNP calls and the software used are correct, is therefore to check out – whenever the corrected tables are published – the samples in Narasimhan et al. (2018) now classified as of R1b1a1a2a1(-) subclades. Or to experiment with the software and available BAM files to see which ones give this result…

6) I don’t know if Khokhlov’s book on Samaran archaeology will contain a reference to the samples, but I doubt it could contribute much more to the genetic data.

The meaning of Yekaterinovka

Of course, the Yekaterinovskiy Cape burials are just a tiny sampling of the dozens of settlements known from Khvalynsk, and the known ones represent just a tiny part of the hundreds that the culture probably had while it developed for more than a thousand years. In that sense, you may say that it is statistically not significant.

Nevertheless, as Anthony’s team recently said when explaining the relevance of their findings at Radzorskoe, the potential implications of any discovery at any of the few studied sites are very important. In this case, by confirming that late Khvalynsk became dominated early by R1b-M269, as was later Yamna, and as were early Yamna offshoots like Afanasevo and Bell Beaker.

I really don’t have anything more to add, whether in comments or per email. That’s as much information and speculation as you can get from me (or from them, I guess). If you want more, you can write to the team members yourselves.

Related: