Admixture of Srubna and Huns in Hungarian conquerors

hungarian-conqueror-migrations

New preprint at BioRxiv, Mitogenomic data indicate admixture components of Asian Hun and Srubnaya origin in the Hungarian Conquerors, by Neparáczki et al. (2018), at BioRxiv.

Abstract (emphasis mine):

It has been widely accepted that the Finno-Ugric Hungarian language, originated from proto Uralic people, was brought into the Carpathian Basin by the Hungarian Conquerors. From the middle of the 19th century this view prevailed against the deep-rooted Hungarian Hun tradition, maintained in folk memory as well as in Hungarian and foreign written medieval sources, which claimed that Hungarians were kinsfolk of the Huns. In order to shed light on the genetic origin of the Conquerors we sequenced 102 mitogenomes from early Conqueror cemeteries and compared them to sequences of all available databases. We applied novel population genetic algorithms, named Shared Haplogroup Distance and MITOMIX, to reveal past admixture of maternal lineages. Phylogenetic and population genetic analysis indicated that more than one third of the Conqueror maternal lineages were derived from Central-Inner Asia and their most probable ultimate sources were the Asian Huns. The rest of the lineages most likely originated from the Bronze Age Potapovka-Poltavka-Srubnaya cultures of the Pontic-Caspian steppe, which area was part of the later European Hun empire. Our data give support to the Hungarian Hun tradition and provides indirect evidence for the genetic connection between Asian and European Huns. Available data imply that the Conquerors did not have a major contribution to the gene pool of the Carpathian Basin, raising doubts about the Conqueror origin of Hungarian language.

hungarian-conqueror-mtdna
“Comparison of major Hg distributions from modern and ancient populations. Asian main Hg-s are designated with brackets. Major Hg distribution of Conqueror samples from this study are very similar to that of other 91 Conquerors taken from previous studies [11,12]. Scythians and ancient Xiongnus show similar Hg composition to the bracketed Asian fraction of the Conqueror samples, but Hg B is present just in Xiongnus. Modern Hungarians have very small Asian components pointing at small contribution from the Conquerors. Of the 289 modern Hungarian mitogenomes 272 are published in [29]. Scythian Hg-s are from [48,49,55,59,71–74]. Xiongnu Hg-s are from [66–69].”

Just recently another article contributed to a similar idea. I already talked about the Bronze Age R1a-z93 sample with high steppe ancestry found in the Balkans, and its likely origin in an expansion of the Srubna or a related culture. No truce, therefore, for those looking for autochthonous continuity anywhere in Europe.

We are seeing how multiple migrations shaped the history of the Carpathian basin (and its complex genetic structure) – and of Europe in general -, often from the Pontic-Caspian steppe. That is clear from many different prehistorical and historical times, such as the expansions of Suvorovo-Novodanilovka, Yamna, Srubna, Thraco-Cimmerians, Sarmatians, Scythians, Huns,…

About the linguistic interpretations based on genetics contained in the paper (Hungarian language as a legacy of Huns), well, you know my stance regarding the Yamnaya ancestral concept (and the wrong linguistic interpretations derived from it, which many sadly keep to this day), and genetics in general to solve language questions

This is yet another example of how (what some people would call) “scientific data” is useless without sound anthropological models.

Featured image, from the article: “Hypothetic origin and migration route of different components of the Hungarian Conquerors. Bluish line frames the Eurasian steppe zone, within which all presumptive ancestors of the Conquerors were found. Yellow area designates the Xiongnu Empire at its zenith from which area the East Eurasian lineages originated. Phylogeographical distribution of modern East Eurasian sequence matches (Fig. 1) well correspond to this territory, especially considering that Yakuts, Evenks and Evens lived more south in the past [108], and European Tatars also originated from this area. Regions where Asian and European Scythian remains were found are labeled green, pink is the presumptive range of the Srubnaya culture. Migrants of Xiongnu origin most likely incorporated descendants of these groups. The map was created using QGIS 2.18.4[109]”.

Article available under a CC-BY-NC-ND 4.0 International license.

Discovered via Razib Khan.

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The Great Hungarian Plain in a time of change in the Balkans – Neolithic, Chalcolithic, and Bronze Age

hungary-yamna-burials-map

I wrote recently about Anthony’s new model of Corded Ware culture expansion from Yamna settlements of Hungary. I am extremely sceptic about it in terms of current genetic finds, and suspicious of the real reasons behind it – probably misinterpretations of the so-called ‘Yamnaya ancestral component’ in recent genetic papers, rather than archaeological finds.

Nevertheless, it means a definitive rejection by Anthony of:

  • The multiple patron-client relationships he proposed to justify a cultural diffusion of Late Indo-European dialects from Yamna into different Corded Ware cultures in the forest-steppe and Forest Zone (see one of his latest summaries of the model in 2015). Now the language change is explained as a pure migration event, and cultural diffusion is not an option. Ergo, if no migration is found from Hungarian Yamna into Lesser Poland, then Corded Ware cultures were not Indo-European-speaking.
  • Ringe’s glottochronological tree for Proto-Indo-European languages (Ringe, Warnow, and Taylor 2002). An early and sudden split of Late PIE dialects in all directions is substituted by a common, Old European language that expanded from a very small area of settlers, in the Carpathian Basin. This is coincident with the current view on North-West Indo-European, and I think that his final acceptance of a sound linguistic model is essential to solve Indo-European questions.
  • The simplistic assumption of Yamna -> Corded Ware -> Bell Beaker migration found in genetic papers of 2015. The new model implies Yamna->Yamna settlers (Eastern Hungary). Yamna settlers are known to have developed into East Bell Beakers (as described by Gimbutas and accepted by Anthony originally, and now also found in the adoption of Heyd’s theory for his new model); therefore a Yamna settlers (Hungary) -> East Bell Beaker evolution is evident and mainstream, now clear also in genetics. It remains to be seen if the additional Yamna settlers (Hungary) -> Proto-Corded Ware migration proposed by him as a novelty in this new model is also right, i.e. if Yamna settlers from Hungary did in fact migrate into sites of Lesser Poland (to form a Proto-Corded Ware culture). If not, then only Heyd’s model remains.

This new model offers thus a more suitable time frame for usual proto-language guesstimates, that would be compatible with a spread of Late Indo-European with Yamna settlers (of R1b lineages) from the steppe into a small region, where North-West Indo-European would have been spoken, and then a potential cultural diffusion through (or founder effect in) a Proto-Corded Ware culture (of R1a-M417 subclades) of Lesser Poland, which is compatible with the Corded Ware Substrate hypothesis.

Since Anthony has stuck his neck out in favour of this new theory – changing some of his popular theories, and rejecting what many geneticists seem to take as certain – , and because of his previous impressive improvements over Gimbutas’ simple steppe theory (now apparently fashionable again), I think he deserves that his proposal of Yamna/Late Indo-European expansion in the Balkans be further investigated, if only to be improved upon.

I recently found the paper 4000-2000 BC in Hungary: The Age of Transformation, by T. Horváth, in Annales Universitatis Apulensis. Series Historica, 20/II, 51-113. While it deals mainly with the potential survival of the Baden culture into the late third millennium BC, it gives some interesting quite early dates for Yamna (‘Pit’) graves in the Carpathian Basin, and potential cultural (and population) movements within the Balkans.

A note about the Corded Ware culture in the Carpathian Basin:

Many researchers may assume that it is unnecessary for us to deal with the Corded Ware and Globular Amphorae cultures of north Germany, Poland and Denmark, and if so it does not matter what the names of the periods are. It actually matters a lot. It is true that in these areas there was no Baden complex, but the period had many Baden (and other) culture “period phenomena”. These seem to part of a larger formation than cultures – evidenced by traces such as cattle burials, the relationship between copper metallurgies and jade – which link these territories even when the culture complexes were different, because these phenomena appear not just in the Baden, but in the Corded Ware and Globular Amphorae area as well (and these cultural complexes partly overlapped each other both in space and time!). Even the characteristics of sites show many similarities: e.g. in the northern part of corded ware distribution area, mainly burials have been discovered (similarly to the Pit Grave culture in the Great Hungarian Plain) and in the southern part only settlements appear.

At the moment we have no explanation regarding the nature of the relationship between them (it is supposed that as a result of geographical conditions the people of the same culture lived in different ecological conditions and they adapted differently to their environment). In considering the whole of Europe around 3500-3000 BC, easily observable settlement signs disappeared (Milisauskas and Kruk, “Late Neolithic/Late Copper Age,” 307), similarly to Hungary, even though in Hungary this occurred from the end of the Middle Copper Age to the Early Bronze Age, between 4000 and 2000 BC. If we do not take into account that the cattle burials of the Baden culture between 3600 and 2800 BC, and possibly even longer than that, have analogies with the cattle burials of areas in the Early and Middle Neolithic Corded Ware culture (because “logically” analogies would be sought in those areas in the Bronze Age but this period is not analogous with that period in those areas), we would not find any spiritual resemblance in their relationships that lies behind their spatial and temporal analogies; cf. comp. Niels Johannsen and Steffen Laursen, “Routes and Wheeled Transport in Late 4th-Early 3rd Millennium Funerary Customs of the Jutland Peninsula: Regional Evidence and European Context,” PZ 85 (2010): 15-58; Horváth “The Intercultural Connections of the Baden „Culture,” 118. It is painful to think about how many relationships we have not explored or even assessed yet!

hungary-yamna-corded-ware-map
One version from both maps shown in the article, by T. Horváth: “Since the two cultures surely lived together in the Late Copper Age, their collective map represents the Late Copper Age (supplemented with Vučedol sites). Since the direction of diffusion of the Kostolac ceramic style is still unclear, two map versions were made. On one the Kostolac followed the Danube River, on the other they diffused in the opposite direction. In northeast Hungary, Coțofeni III appeared. On this map Kostolac sites are not depicted as dots but, in light of their position and density, proportionately sized arrows are used.”

On Yamna culture and burials in the Carpathian Basin:

Looking at Pit Grave kurgans on the distribution map, it is apparent that burials are the densest where there were no Boleráz or Baden occupations (in this respect this was a kind of “no man’s land”, but from the whole Late Copper Age perspective it was not: the sites of the Baden complex and Pit Grave complemented each other and even partially overlapped). Apart from burials, no Pit Grave settlements or other types of Pit Grave sites are known in Hungary, therefore we do not know whether Pit Grave settlements were situated near the kurgans or whether were somewhere else entirely and we simply have not found them yet.

Since the Pit Grave people had a different lifestyle from the Baden, we can assume that, up to the line of the Tisza River, small animal-keeping mobile groups (Pit Grave) met more populated and settled, agriculturalist, indigenous Boleráz-Baden groups. Animal keepers (Pit Grave) settled in areas where agriculturalists (Boleráz and Baden) did not; in some places, however, they crossed each other’s paths (Fig. 5, 7). Sometimes their sites are very close to each other, sometimes they appear on one site and they can be identified in the stratigraphy of a site. In the latter case the kurgan is always situated on top of a Baden settlement, indicating that Pit Grave not only followed the Baden at these sites but may have represented a somewhat higher social power and belief system than the Baden.

The relationship between pastoral, patrilineal, combatant nomadic tribes and agriculturalist communities is often described as some sort of patron and client relationship. In reality, the signs of such assumption are not visible in the Pit Grave-Baden relationship. There are cases when more aggressive herders conquered more developed agriculturalist communities, but there are also cases when the conqueror’s culture was more developed or stronger than that of the conquered. Always, the conquering nomads are the patrons, the rulers and the empire builders.

In our case, timing is important. How much time had passed on those common sites where a Baden settlement was followed by a Pit Grave kurgan? In these cases, it is certain that the kurgan is younger, but how much younger?

hungary-yamna-settlements
From the article, by T. Horváth. “On the 10 locations analysed, surviving Baden can be assumed after 2800 BC. Unfortunately, it is not possible to predict which sites would survive further scrutiny of radiocarbon dating in this respect; only a few dates are available so far. Therefore, on the map of Baden that still existing after the Late Copper Age, I have also represented all sites (up to the Danube River line) and combined them with Early Bronze Age sites. Since the majority of Makó sites are represented by only one find (scattered finds), and the majority of sites have just one grave, it is impossible to ascertain whether it was part of a cemetery, was within a settlement, or was an individual burial without any further features. Therefore, following Dani 2005, I utilized subdivisions: perhaps in the future this fine subdivision will provide a meaningful explanation (1). Since the radiocarbon dates of Pit Grave kurgans clearly show that the Pit Grave survived at least until 2500 BC, I combined the previous map with that of the Pit Grave. This map would show a realistic picture of cultures after 2800 BC east of the Danube River (2).”

To sum up, the Pit Grave and Baden in the Late Copper Age were certainly contemporary from 3350 BC in the Great Hungarian Plain, and they had common sites, sites which were very close to each other, sites which were far from each other, and also independent sites. The Pit Grave culture surely survived in the transitional period, and into Early Bronze Age I, but perhaps even longer. For the most part, the Baden had ended by 2900 BC in the Great Hungarian Plain. Mapping and some other data (e.g. the discovery that Younger-type, not Mondsee-type, metal objects, which can now be considered to be Baden, even appear east of the Danube River) does not exclude the possibility of searching further for traces of Baden surviving in the Great Hungarian Plain together with or alongside to the Pit Grave. On the common Baden-Pit Grave sites, even without carbon dating, we can assume from already known stratigraphical data that they closely followed each other in time.

For those of you interested in more detailed radiocarbon analysis and assessment of Yamna burials and settlements, from the steppe to the Balkans, to investigate Anthony’s theory further – apart from those authors referenced by him – , I can recommend reading Y. Rassamakin (e.g. Import and Imitation in Archaeology, 2008), S. Ivanova, or Claudia Gerling (e.g. Prehistoric Mobility and Diet in the West Eurasian Steppes 3500 to 300 BC).

Featured image, from the article, by T. Horváth: Distribution map of the Pit Grave.

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Genetic landscapes showing human genetic diversity aligning with geography

world-effective-migration

New preprint at BioRxiv, Genetic landscapes reveal how human genetic diversity aligns with geography, by Peter, Petkova, and Novembre (2017).

Abstract:

Summarizing spatial patterns in human genetic diversity to understand population history has been a persistent goal for human geneticists. Here, we use a recently developed spatially explicit method to estimate “effective migration” surfaces to visualize how human genetic diversity is geographically structured (the EEMS method). The resulting surfaces are “rugged”, which indicates the relationship between genetic and geographic distance is heterogenous and distorted as a rule. Most prominently, topographic and marine features regularly align with increased genetic differentiation (e.g. the Sahara desert, Mediterranean Sea or Himalaya at large scales; the Adriatic, inter-island straits in near Oceania at smaller scales). We also see traces of historical migrations and boundaries of language families. These results provide visualizations of human genetic diversity that reveal local patterns of differentiation in detail and emphasize that while genetic similarity generally decays with geographic distance, there have regularly been factors that subtly distort the underlying relationship across space observed today. The fine-scale population structure depicted here is relevant to understanding complex processes of human population history and may provide insights for geographic patterning in rare variants and heritable disease risk.

world-migration-effective
Regional patterns of genetic diversity. a: scale bar for relative effective migration rate. Posterior effective migration surfaces for b: Western Eurasia (WEA) e: Central/Eastern Eurasia (CEA) g: Africa (AFR) h Southern African hunter-gatherers (SAHG) k: and Southeast Asian (SEA) analysis panels. ‘X’ marks locations of samples noted as displaced or recently admixed, ‘H’ denotes Hunter-Gatherer populations (both ‘X’ and ‘H’ samples are omitted from the EEMS model fit); in panel g, red circles indicate Nilo-Saharan speakers and in panel h, ‘B’ denotes Bantu-speaking populations. Approximate location of troughs are shown with dashed lines (see Extended Data Figure 4). PCA plots: c: WEA d:Europeans in WEA f: CEA i: SAHG j: AFR l: SEA. Individuals are displayed as grey dots. Large dots reflect median PC position for a sample; with colors reflecting geography matched to the corresponding EEMS figure. In the EEMS plots, approximate sample locations are annotated. For exact locations, see annotated Extended Data Figure 4 and Table S1. Features discussed in the main text and supplement are labeled. FST values per panelemphasize the low absolute levels of differentiation.”

Among ‘effective migration surfaces‘ (or potential past migration routes), the Pontic-Caspian steppe and its most direct connection with the Carpathian basin, the Danubian plains, appear maybe paradoxically as a constant ‘trough’ (below average migration rate) in all maps.

After all, we could have agreed that this region should be a priori thought as the route of many migrations from the steppe and Asia into Central Europe (and thus of ‘effective migration’) in prehistoric, proto-historic and historic times, such as Suvorovo-Novodanilovka (Pre-Anatolian), Yamna (Late Indo-European), probably Srubna, Scythian-Cimmerian, Sarmatian, Huns, Goths, Avars, Slavs, Mongols

It most likely (at least partially) represents a rather recent historical barrier to admixture, involving successive Byzantine, South Slavic, and Ottoman spheres of influence positioned against Balto-Slavic societies of Eastern Europe.

europe-migration-routes
Location of troughs in West Eurasia (below average migration rate in more than 95% of MCMC iterations) are given in brown. Sample locations and EEMS grid are displayed for the West Eurasian analysis panel. FST values are provided per panel to emphasize the low absolute levels of differentiation.

Featured image, from the article: “Large-scale patterns of population structure. a: EEMS posterior mean effective migration surface for Afro-Eurasia (AEA) panel. ‘X’ marks locations of samples excluded as displaced or recently admixed. ‘H marks locations of excluded hunter-gatherer populations. Regions and features discussed in the main text are labeled. Approximate locations of troughs are annotated with dashed lines (see Extended Data Figure 4). b: PCA plot of AEA panel: Individuals are displayed as grey dots, colored dots reflect median of sample locations; with colors reflecting geography and matching with the EEMS plot. Locations displayed in the EEMS plot reflect the position of populations after alignment to grid vertices used in the model (see methods).”

Images and text available under a CC-BY-NC-ND 4.0 International License.

Discovered via Razib Khan’s blog.

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