Preprint of a review by Iosif Lazaridis, The evolutionary history of human populations in Europe.
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Steppe populations during the Eneolithic to Bronze Age were a mix of at least two elements, the EHG who lived in eastern Europe ~8kya and a southern population element related to present-day Armenians, and ancient Caucasus hunter-gatherers, and farmers from Iran. Steppe migrants made a massive impact in Central and Northern Europe post- 5kya[28,43]. Some of them expanded eastward, founding the Afanasievo culture and also eventually reached India. These expansions are probable vectors for the spread of Late Proto-Indo-European languages